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Marisora unimarginata (COPE, 1862)

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Higher TaxaScincidae, Mabuyinae, Sauria (lizards) 
Subspecies 
Common NamesCentral American Mabuya 
SynonymMabuya unimarginata COPE 1862: 187
Mabuya unimarginata — TAYLOR 1956: 302
Mabuya unimarginata — LINER 1994
Mabuya unimarginata — GREER & NUSSBAUM 2000
Mabuya unimarginata — KÖHLER 2000: 92
Mabuya unimarginata — MAUSFELD et al. 2002
Marisora unimarginata — HEDGES & CONN 2012: 119 
DistributionW Costa Rica, Panama

Type locality: “Panama” Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
TypesHolotype: unlocated and presumably lost 
CommentSynonymy: Mabuya unimarginata was listed as synonym of Mabuia aurata in BOULENGER 1887: 189.

Reproduction: viviparous.

Type Species: Mabuya unimarginata Cope, 1862:187 is the type species of the genus Marisora HEDGES & CONN 2012.

Diagnosis (genus). Species of the Genus Marisora are characterized by (1) frontoparietals, two (rarely three), (2) supraciliaries, four (occasionally three, five, or six), (3) supraoculars, four (rarely three), (4) prefrontal contact, absent (or contact very rarely), (5) parietal contact, present (or occasionally no contact), (6) rows of nuchals, one (rarely two rows), (7) dorsals + ventrals, 109–131, (8) total lamellae, 184–229, (9) a dark middorsal stripe, absent, (10) dark dorsolateral stripes, usually absent (present in M. alliacea comb. nov.), (11) a dark lateral stripe, present, and (12) dark ventral striping, absent. Species of Marisora are medium to large, with a range of maximum body sizes among the species of 82–95 mm SVL (except for one species, M. magnacornae sp. nov., known from a single 77.4 mm specimen; Table 2).
All Marisora have a basic pattern, usually bold and well-defined, of a dark lateral band several scales wide bordered below by a narrow pale stripe usually less than one scale wide. In two species (M. aurulae sp. nov. and M. falconensis), this basic pattern is weakly defined, and in another (M. alliacea), there are additional (dorsolateral) dark stripes. The absence of dark dorsolateral stripes (except in M. alliacea) distinguishes this genus from Aspronema, Brasiliscincus (most individuals), Manciola, Orosaura, Panopa, Psychosaura, Spondylurus, and Varzea (most individuals). The presence of one row of nuchals (rarely two) distinguishes the Genus Marisora from Exila and Panopa (2–5 rows) and most Spondylurus (usually 2 rows). The presence of two (rarely three) frontoparietals (instead of one fused scale) distinguishes this genus from Exila, Notomabuya, and Panopa. The presence of a pale lateral stripe and absence of dark ventral striping distinguish this genus from the Genus Alinea. The absence of a middorsal dark stripe further distinguishes this genus from Aspronema. The presence of four (usually) supraciliaries (versus 5–6) distinguishes Marisora from Capitellum and Exila. Contact (usually) of the parietal scales distinguishes this genus from the Genus Copeoglossum (parietals usually not in contact). In having four supraoculars (rarely three), Marisora is separated from two genera with three supraoculars: Aspronema (rarely four) and Mabuya (rarely two or four). In having 184–229 total lamellae, it is distinguished from Manciola (147– 154 lamellae) and Alinea (231–259 total lamellae). From Maracaiba, it differs (weakly) by having a low number of dorsals (50–63 versus 63; only three of 80 Marisora with 63 dorsals) [HEDGES & CONN 2012]. 
EtymologyThe generic name (Marisora) is a feminine noun derived from the Latin words maris (sea) and ora (coast, or border), referring to the distribution of this genus occurring predominately in low elevations near the coast (Caribbean, Atlantic, and Pacific), with relatively few inland and upland localities. Three of the seven species occur exclusively on islands. 
References
  • Boulenger, G. A. 1887. Catalogue of the Lizards in the British Museum (Nat. Hist.) III. Lacertidae, Gerrhosauridae, Scincidae, Anelytropsidae, Dibamidae, Chamaeleontidae. London: 575pp. - get paper here
  • CASTIGLIA,RICCARDO; FLAVIA ANNESI, ALEXANDRA M.R. BEZERRA, ANDRÉS GARCÍA & OSCAR FLORES-VILLELA 2010. Cytotaxonomy and DNA taxonomy of lizards (Squamata, Sauria) from a tropical dry forest in the Chamela-Cuixmala Biosphere Reserve on the coast of Jalisco, Mexico. Zootaxa 2508: 1–29 - get paper here
  • Cope, E.D. 1863. Contributions to Neotropical saurology. Proc. Acad. Nat. Sci. Philadelphia 14 [1862]: 176-188. - get paper here
  • Cope, E.D. 1876. On the Batrachia and Reptilia of Costa Rica with notes on the herpetology and ichthyology of Nicaragua and Peru. Journal of the Academy of Natural Sciences of Philadelphia N.S. (2) 8: 93-183 [1875] - get paper here
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  • Lee, J.C. 1996. The amphibians and reptiles of the Yucatán Peninsula. Comstock, Cornell University Press, Ithaca, 500 pp.
  • Mausfeld, Patrick; Andreas SCHMITZ; Wolfgang BÖHME; Bernhard MISOF; Davor VRCIBRADIC; Carlos Freder 2002. Phylogenetic Affinities of Mabuya atlantica Schmidt, 1945, Endemic to the Atlantic Ocean Archipelago of Fernando de Noronha (Brazil): Necessity of Partitioning the Genus Mabuya Fitzinger, 1826 (Scincidae: Lygosominae). Zool. Anz. 241: 281–293
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  • Mendenhall, Chase D.; Luke O. Frishkoff, Georgina Santos-Barrera, Jesús Pacheco, Eyobed Mesfun, Fern 2014. Countryside biogeography of Neotropical reptiles and amphibians. Ecology 95:856–870, http://dx.doi.org/10.1890/12-2017.1 - get paper here
  • Mendoza Quijano, F. et al. 2002. Mabuya unimarginata COPE 1862. Mexico: Hidalgo... Elaphe 10 (4): 68
  • Miralles, Aurélien 2006. A new species of Mabuya (Reptilia, Squamata, Scincidae) from the Caribbean Island of San Andrés, with a new interpretation of nuchal scales: a character of taxonomic importance. The Herpetological Journal 16 (1):1-7.
  • Savage, J.M. 1973. Herpetofauna of Costa Rica.
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