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Nactus intrudusus ZUG, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Markham Slender-toed Gecko 
SynonymNactus intrudusus ZUG 2020: 47 
DistributionPapua New Guinea (Markham River valley, Morobe Province)

Type locality: Papua New Guinea, Morobe Province, Busu River, 8 miles [~12.9 km] north of Lae [−6.7333° 146.9999°].  
TypesHolotype. AMNH 95175, adult male, collected by R. G. Zweifel and G. Sluder on 14 August 1964 (Figure 19).
Paratypes. All from Papua New Guinea, Morobe Province. Adult females: AMNH 92664, Upper Markham Valley, Umi River (?), 475 m collected by H. M. Van Duesen on 17 November 1959; AMNH 95177, Lae [−6.7333° 146.9999°], collected by R. G. Zweifel on 17 July 1964; AMNH 103243, 7 miles [~11.3 km] (road) north of Lae [~−6.6352 146.9808], collected by R. G. Zweifel on 25 August 1968. Adult male: AMNH 103242, 7 miles [~11.3 km] (road) north of Lae [~−6.6352 146.9808], collected by R. G. Zweifel on 25 August 1968. 
DiagnosisDefinition. A bisexual taxon of geckos (Gek- koninae) with moderate to large adults (♀♀ 53–65 mm, ♂♂ 55–65 mm SVL) with smooth subcaudals, moderate to large postmental scales (usually ≤10), no or slight postmental- infralabial (0–1) contact, moderate number of dorsal tubercle rows (14–15) and moderately high number of tubercles (26–36) in parasagittal row (TubRow), no enlarged tubercles on dorsal surface of hindlimb, and modest number of precloacal pores (9–11) in males and none in females. Diagnostic summary in Table 8.

Coloration in life not reported.

Variation. Females and males display only slight metric differences (adult ♀♀ 59.2, 52.7–65.0 mm; adult ♂♂58.4, 55.3–63.0 mm SVL) and share similar proportions (total adult sample): TrunkL/SVL 42.0%, 36%–46%; HeadL/ SVL 26.7%, 26%–27%; HeadW/SVL 17.9%, 17%–19%; HeadW/HeadL 66.9%, 62%–69%; EyeD/SVL 7.2%, 6–8%; EyeD/HeadL 27.0%, 24%–29%; NarEye/HeadL 32.1%, 31%–33%; Interorb/HeadL 32.9%, 29%–37%; SnW/HeadL 16.2%, 15–17%.
Typical head scale pattern of small, granular scales except for those bordering nares and mouth. Snout with large ros- tral scale bearing middorsal cleft posteriorly; large supral- abials (4, 3–4) and infralabials (4, 3–4) in front of anterior edge of orbit, first supra- and infralabial largest of respective series. Chin with large trapezoidal mental and usually mod- erately large postmentals (Postm = 10, 2–20) and usually no genial scale between postmental and first supralabial (0, 0–1 PmLab); dorsally, trunk with 14, 14–15 (DorsTub) enlarged tubercles transversely on dorsum and with 29, 26–36 tubercles (TubRow) longitudinally along trunk; 8, 7–9 tubercle rows transversely between hindlimbs (TubHip). Uniform cover- ing of small tuberculate-like scale on dorsal half of forelimb and hindlimb (TubHindl = 0). Tail with small uniform scales dorsally and laterally, ventrally with uniform, small usually smooth scales (Subcaud = 1, 0–1). Usually, single cloacal spurs (1, 1–2 CloacS) on each side. Ventral scales from chin to vent small, granular, somewhat larger on chest and abdomen. Pre- cloacal pores, males 9–11, females 0. Forefoot with 17, 17–18 lamellae beneath fourth digit (4FingLm), 6, 5–6 scale rows on palm; hindfoot with 20, 20–22 lamellae beneath fourth toe (4ToeLm). 
CommentDistribution: see map in Zug 2020: 47 (Fig. 18). 
EtymologyThis unique population seemingly intrudes between north coast populations with tuberculate limbs and keeled subcaudal scales, hence the use of the Latin intrudo, intrudusus for “intruder.” The specific name is proposed as a noun in apposition. 
  • Zug, George R. 2020. Diversity in Pacific Slender-Toed Geckos, Nactus pelagicus Complex (Reptilia: Squamata), of New Guinea and Adjacent Islands. Smithsonian Contributions to Zoology, (651): x + 92 pages - get paper here
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