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Naja sumatrana MÜLLER, 1890

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Higher TaxaElapidae, Serpentes (snakes) 
Subspecies 
Common NamesE: Equatorial Spitting cobra, Golden Spitting Cobra
G: Äquatoriale Speikobra, Sumatra-Kobra 
SynonymNaja tripudians var. sumatrana MÜLLER 1890: 277
Naja tripudians var. nigra PETERS 1859: 271 (nomen neglecta)
Naia tripudians var leucodira BOULENGER 1896
Naia tripudians var. miolepis BOULENGER 1896
Naia tripudians var. caeca BOULENGER 1896 (part.)
Naia tripudians var. sputatrix — BOULENGER (part.) 1896
Naja sumatrana miolepis BOULENGER 1896
Naja tripudians paucisquamis PERACCA 1899
Naja tripudians sumatrana — SCHENCKEL 1901
Naia tripudians leucodira ROSEN 1905
Naja naja sputatrix — STEJNEGER (part) 1907
Naja naja miolepis GRIFFIN 1909
Naja naja paucisquamis BARBOUR 1912
Naja naja leucodira MERTENS 1934
Naja naja sumatrana — KOPSTEIN 1935
Naja kaouthia sputatrix — DERANIYAGALA 1960
Naja sputatrix malayae DERANIYAGALA 1961
Naja sputatrix miolepis DERANIYAGALA 1961
Naja naja sumatrana — HARDING & WELCH 1980
Naja sputatrix sputatrix — LINGENHOLE & TRUTNAU 1989
Naja sumatrana — WÜSTER & THORPE 1989
Naja sumatrana — MANTHEY & GROSSMANN 1997: 424
Naja sumatrana — COX et al. 1998: 29
Naja (Naja) sumatrana — WALLACH et al. 2009 
DistributionMalaysia (Peninsula, East Malaysia, Pulau Tioman), Thailand, Singapore, Indonesia (Sumatra, Borneo, Bangka, Belitung),
Philippines (Palawan: Calamian Islands etc.)

Type locality: Solok, Sumatra. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
TypesHolotype, NMBA 2244, a female (F. Müller). 
CommentVenomous!

Populations from the Malayan Peninsula were for a long time referred to as Naja naja sputatrix, those from Borneo as Naja naja miolepis.

N. tripudans var. nigra: type: ZMB 7098, terra typica: Sarawak Borneo Indonesia, leg Doria.

Reproduction: oviparous. 
References
  • Auliya, M. 2006. Taxonomy, Life History, and conservation of giant reptiles in west Kalimantan. Natur und Tier Verlag, Münster, 432 pp. - get paper here
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
  • Gaulke M. 1999. Die Herpetofauna von Calauit Island (Calamianes-Inseln, Provinz Palawan, Philippinen) (Amphibia et Reptilia). Faun. Abh. Staatl. Mus. Tierk. Dresden 21 (19)
  • Grismer, L.L., McGuire, J.A.; Sosa, R. & Kaiser, H. 2002. Revised checklist and comments on the terrestrial herpetofauna of Pulau Tioman, Peninsular Malaysia. Herpetological Review 33 (1): 26-29 - get paper here
  • Grossmann,W. & Tillack,F. 2001. Bemerkungen zur Herpetofauna des Khao Lak, Phang Nga, thailändische Halbinsel. Teil II: Reptilia: Serpentes; Testudines; Diskussion. Sauria 23 (1): 25-40 - get paper here
  • Grossmann,W. & Tillack,F. 2001. Bemerkungen zur Herpetofauna des Khao Lak, Phang Nga, thailändische Halbinsel. Teil III: Ergebnisse der Jahre 1999 und 2000. Sauria 23 (3): 21-34 - get paper here
  • Kopstein, Felix 1935. Herpetologische Notizen XI. Bemerkung über Naja naja leucodira (Blgr.) Treubia 15 (2): 151-155
  • Leviton,A.E. 1965. Contributions to a review of Philippine snakes. VII. The snakes of the genera Naja and Ophiophagus. Philippine J. Sci. 93: 531-550 - get paper here
  • Lingenhöle, S. & Trutnau, L. 1989. Über die Kobras der Gattung Naja LAURENTI 1758 in Thailand. Herpetofauna 11 (58): 6-13 - get paper here
  • Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Müller,F. 1887. Fünfter Nachtrag zum Katalog der herpetologischen Sammlung des Basler Museums. Verh. Naturf. Ges. Basel 8: 249-296 - get paper here
  • Peracca, M.G. 1899. Voyage de MM. M. Bedot et C. Pictet dans l'Archipel Malais. Reptiles et Batraciens de l'Archipel Malais. Revue suisse de Zoologie. Annales de la Societe Zoologique suisse et du Musee d'Histoire naturelle de Geneve 7(2): 321-330 - get paper here
  • Peters, Wilhem Carl Hartwig 1859. Nachrichten von Hrn. Fedor Jagor, der auf eigne Kosten nach Ostindien und den Philippinen gereist ist und von der Akademie mit Instructionen für das Sammeln naturwissenschaftlicher Gegenstände versehen wurde. Monatsber. königl. Akad. Wiss. Berlin. 1859 (März): 269-271 - get paper here
  • Rooijen, J. 2009. Estimating the snake species richness of the Santubong Peninsula (Borneo) in two different ways. Contributions to Zoology 78 (4): 141-147
  • Slowinski, Joseph B. and Wolfgang Wüster. 2000. A new cobra (Elapidae: Naja) from Myanmar (Burma). Herpetologica 56 (2): 257-270 - get paper here
  • Teo, R.C.H. & Rajathurai, S. 1997. Mammals, reptiles and amphibians in the Nature Reserves of Singapore - diversity, abundance and distribution. Proc. Nature Reserves Survey Seminar. Gardens’ Bulletin Singapore 49: 353-425
  • van Rooijen, Johan and Myriam van Rooijen. 2007. The land snakes of the Santubong Peninsula, Sarawak, Borneo: A preliminary list of species with natural history notes. Russian Journal of Herpetology 14 (1):27-38 - get paper here
  • Van Rooijen, Johan, Chan Kin Onn, L. Lee Grismer & Norhayati Ahmad 2011. Estimating the herpetofaunal species richness of Pangkor Island, Peninsular Malaysia. Bonn zoological Bulletin 60 (1): 3-8 - get paper here
  • Wallach, V.; Wüster, W. & Broadley, D.G. 2009. In praise of subgenera: taxonomic status of cobras of the genus Naja Laurenti (Serpentes: Elapidae). Zootaxa 2236: 26–36 - get paper here
  • Wüster, Wolfgang 1993. A century of confusion: Asiatic cobras revisited Vivarium 4 (4): 14-18
  • Wüster,W. 1996. Taxonomic changes and toxinology: Systematic revisions of the asiatic cobras (Naja naja complex). Toxicon 34 (4): 399-406
  • Wüster,W.; Thorpe,R.S. 1989. Population affinities of the Asiatic cobra (Naja naja) species complex in south-east Asia: reliability and random resampling. Biol. J. Linn. Soc. 36: 391-409. - get paper here
  • Wüster,W.; Thorpe,R.S. 1991. Asiatic cobras: Systematics and snakebite. Experientia 47: 205-209
 
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