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Ninia schmidti (JAN, 1862)

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Schmidt’s Coffee-Snake.
Spanish: Culebra cafetera de Schmidt 
SynonymStreptophorus sebae schmidti JAN 1862: 27
Ninia spilogaster PETERS 1881: 49
Ninia schmidti — ARTEAGA & HARRIS 2023: 326 
DistributionE Panama, Colombia (incl. Valle del Cauca), Ecuador, Venezuela (Táchira, Barinas [HR 35: 192]), Trinidad

Type locality: “Guayaquil”  
Reproductionoviparous 
TypesNeotype. ZMH R10390 (Fig. 9), adult female collected by Bi. Jansen between 1901 and 1902 at Guayaquil, Guayas province, Ecuador. Holotype: lost, was ZMH [schmidti]
Holotype: unlocated at ZMB fide Arteaga & Harris 2023 [spilogaster] 
DiagnosisDiagnosis. Ninia schmidti comb. nov. is placed in the genus Ninia, as diagnosed by Dunn (1935), based on phylogenetic evidence (Fig. 1). The species is diagnosed based on the following combination of characters: (1) 19/19/19 keeled dorsals; (2) two postoculars; (3) loreal 1.4–2.3 × longer than high; (4) temporals 1+2; (5) seven supralabials, third and fourth contacting orbit; (6) seven or eight infralabials, first four or five contacting chin shields; (7) usually two rows of chin shields; (8) one or two preventrals; (9) 138–144 ventrals in males, 139–155 in females; (10) 50–57 subcaudals in males, 46–53 in females; (11) dorsal ground color uniformly black without a white nuchal collar (Fig. 10); (12) ventral surfaces of adults obscured with dark pigment particularly along the posterior edge of each ventral scale (Figs 9b, 11a), immaculate white in some juveniles (Fig. 11b); (13) 167–283 mm SVL in males, 230–409 mm in females; (14) 42–61 mm CL in males, 53–84 mm in females. (Arteaga & Harris 2023)


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CommentSynonymy: previously a synonym of N. atrata, but revalidated by Arteaga & Harris 2023. 
EtymologyNamed after Philipp Moses Paul Frederich Schmidt (1800–1869/1873), a physician in Hamburg best known for his work on sea snakes (Beolens et al. 2011). 
References
  • Arteaga, Alejandro and Kyle J. Harris 2023. A New Species of Ninia (Serpentes, Colubridae) from western Ecuador and Revalidation of N. schmidti. Evolutionary Systematics. 7(2): 317-334 - get paper here
  • Jan, G. 1862. Enumerazione sistematico delle specie d'ofidi del gruppo Calamaridae. Arch. Zool., Anat. Fisiol., 2: 1-76. - get paper here
 
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