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Nucras boulengeri NEUMANN, 1900

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Higher TaxaLacertidae, Eremiadinae, Sauria, Lacertoidea, Squamata (lizards)
Subspecies 
Common NamesE: Uganda Savannah Lizard, Boulenger’s scrub lizard 
SynonymNucras boulengeri NEUMANN 1900: 56
Nucras emini BOULENGER 1907 (fide LOVERIDGE 1957: 230)
Nucras ukerewensis BOLKAY 1909 (fide BISCHOFF 2024)
Nucras boulengeri — BOULENGER 1920: 16
Nucras emini — BOULENGER 1920: 10
Nucras kilosae LOVERIDGE 1922: 314 (fide BROADLEY & HOWELL 1991)
Nucras emini — LOVERIDGE 1923
Nucras boulengeri boulengeri — LOVERIDGE 1955: 176
Nucras boulengeri boulengeri — LOVERIDGE 1957: 7
Nucras boulengeri — BROADLEY & HOWELL 1991: 17
Nucras boulengeri kilosae — BISCHOFF 1991
Nucras boulengeri kilosae — BAUER & GÜNTHER 1995
Nucras boulengeri — HAAGNER et al. 2000
Nucras boulengeri — SPAWLS et al 2018: 192
Nucras boulengeri — BAUER et al. 2025 
DistributionUganda, N Tanzania, Kenya, E/C Tanzania, Zambia

Type locality: Lubwa’s = Luba’s village, Busoga, Uganda.

Type locality: Tindiga, 5 miles south of Kilosa, Morogoro Dist., Tanzania [kilosae]

Type locality: Shirati, Tansania, east bank of Lake Victoria (Victoria Nyanza) [ukerewensis]  
Reproductionoviparous 
TypesType: BMNH 1946.8.6.16 (and possibly additional specimens).
Syntypes: BMNH [kilosae]
Holotype: lost during the Russian attack on Budapest 1956, until then in the Hungarian National Museum [ukerewensis] 
DiagnosisAdditional details, e.g. a detailed description or comparisons (1159 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSynonymy: Broadley and Howell (1991) synonymized N. kilosae with N. boulengeri.

Distribution: For localities in Zambia see map in Bauer et al. 2025: 488 (Fig. 5). 
EtymologyNamed after George Albert Boulenger (1858-1937), herpetologist at the British Museum of Natural History, London. For comments on the types of N. b. kilosae see BAUER & GÜNTHER 1995.

Nucras emini was named after Emin Pascha (1840-1892), German researcher, also known as Isaak Eduard Schnitzer (now a synonym)

Nucras ukerewensis was named after Ukerewe, the largest island in Lake Victoria (now a synonym) 
References
  • BAUER, A. M., CHILDERS, J. L., & BURGER, M. 2025. The Nucras (Squamata: Lacertidae) of the eastern Zambezi River Basin, with the description of a new species from Zambia. Zootaxa, 5632(3), 480-500 - get paper here
  • Bauer, Aaron M.;Günther, Rainer 1995. An annotated type catalogue of the lacertids (Reptilia: Lacertidae) in the Zoological Museum, Berlin. Mitt. Zool. Mus. Berlin 71 (1): 37-62 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Bischoff, Wolfgang 2024. Ein „fast vergessenes“ Synonym von Nucras boulengeri Neumann, 1900. Die Eidechse 35 (2): 54-56
  • Bolkay, S. 1909. Description of a new lizard from the Genus Nucras Gray. Archivum Zoo- logicum, Budapest, 1(3): 13-14
  • Boulenger, G.A. 1907. Descriptions of new lizards in the British Museum. Ann. Mag. Nat. Hist. (7) 19: 486-489. - get paper here
  • Boulenger,G.A. 1920. Monograph of the Lacertidae, vol. 1. British Museum (London) - get paper here
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Buruwate, T. C., & Lloyd-Jones, D. J. 2024. Amphibian and Reptile Diversity of Niassa Special Reserve, Northern Mozambique. Journal of East African Natural History, 113(1), 1-18 - get paper here
  • Edwards, Shelley; Krystal A. Tolley, Bieke Vanhooydonck, G. John Measey and Anthony Herrel 2013. Is dietary niche breadth linked to morphology and performance in Sandveld lizards Nucras (Sauria: Lacertidae)? Biological Journal of the Linnean Society, DOI: 10.1111/bij.12148 - get paper here
  • Haagner,G.V.; Branch,W.R. & Haagner,A.J.F. 2000. Notes on a collection of reptiles from Zambia and adjacent areas of the Democratic Republic of the Congo. Annals of the Eastern Cape Museum 1: 1 – 25
  • Loveridge, A. 1923. Notes on East African lizards collected 1920-1923 with the description of two new races of Agama lionotus Blgr. Proc. Zool. Soc. London 1923: 935-969 - get paper here
  • Loveridge, A. 1955. On a second collection of reptiles and amphibians taken in Tanganyika Territory by C. J. P. lonides,. Journ. E. Africa Nat. Hist. Soc. 22: 169-198
  • Loveridge, A. 1957. On a third collection of reptiles taken in Tanganyika by C.J.P. Ionides, Esq. Tanganyika Notes and Records 43: 1-19
  • Loveridge,A. 1922. New reptiles from Tanganyika Territory. Proc. Zool. Soc. London 1922: 313-315 - get paper here
  • Malonza, Patrick K.; David M. Mulwa, Joash O. Nyamache, Georgina Jones 2017. Biogeography of the Shimba Hills ecosystem herpetofauna in Kenya. Zoological Research 38(5): 1-11 - get paper here
  • Muchai, Vincent & Patrick Malonza 2011. The reptiles and amphibians of Kibwezi Forest, Ngulia and Chyulu Hills. Kenya Past and Present (39): 45-49 - get paper here
  • MÜLLER, J. 1998. Zur besseren Kenntnis von Nucras boulengeri (NEUMANN, 1900), einer wenig bekannten Eidechse aus Ostafrika. Die Eidechse 8 (3): 88-96 - get paper here
  • Neumann, O. 1900. Description of a new lizard of the genus Nucras from Usoga, British East Africa. Ann. Mag. Nat. Hist. (7) 5: 56 - get paper here
  • Pietersen, Darren, Verburgt, Luke & Davies, John 2021. Snakes and other reptiles of Zambia and Malawi. Struik Nature / Penguin Random House South Africa, 376 pp., ISBN 9781775847373
  • Razzetti, E. & Msuya, C.A. 2002. Field guide to the amphibians and reptiles of Arusha National Park (Tanzania). Publ. Ed. Negri Istituto, Oikos, Varese, 84 pp.
  • Spawls, S. 2017. A Guide to Amphibians and Reptiles of the Maasai Mara Ecosystem. Norwich City College Print Room - get paper here
  • Spawls, S. & Rotich, D. 1997. An annotated checklist of the lizards of Kenya. J. East African Nat. Hist. 86: 61-83
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
 
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