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Oedura marmorata GRAY, 1842

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Higher TaxaDiplodactylidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesMarbled Velvet Gecko 
SynonymOedura marmorata GRAY 1842
Phyllodactylus (Aedura) marmorata - DUMÉRIL 1856
Phyllodactylus marmoratus - STEINDACHNER 1867
Oedura verrillii COPE 1869: 318
Oedura verrillii - GÜNTHER 1869
Oedura fracticolor DE VIS 1884
Oedura marmorata — BOULENGER 1885: 104
Oedura ? verrillii — BOULENGER 1885: 108
Oedura greeri WELLS & WELLINGTON 1985: 14 (nom. nud.)
Oedura marmorata — RÖSLER 2000: 97
Oedura marmorata — COGGER 2000: 263
Oedura marmorata — OLIVER et al. 2012
Oedura marmorata — OLIVER & DOUGHTY 2016 
DistributionAustralia (Northern Territory)

Type locality: Port Essington, N. T. [lectotype]

Type locality: ‘Kimberley’ (currently Normanton) at the mouth of the Norman River on the Gulf of Carpentaria in Queensland [fracticolor] Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesLectotype BMNH xxii.2b.; syntype: MCZ R-6471
Holotype: AMS R87677 [greeri] 
CommentSynonymy: OLIVER & DOUGHTY 2016 split up O. marmorata into several spcies, and removed Oedura cincta DE VIS 1888 from the synonymy of O. marmorata. KLUGE (1965, 1993) listed verillii with a question mark, indicating its unclear synonymy. Owing to the vague description and type location and lack of type specimens, OLIVER & DOUGHTY 2016 maintain O. verrillii as a junior synonym of O. marmorata. Oedura greeri Wells & Wellington, 1985 (holotype: AMS R87677) was described without diagnosis and is regarded as a nomen nudum (Shea & Sadlier 1999).

Type species: Oedura marmorata is the type species of the genus Oedura GRAY 1842. Cogger (1992) offers the following generic diagnosis: “Rostral and mental shields rounded. Labials larger than adjacent scales. Postmentals enlarged. Digits moderately long,depressed, moderately expanded distally to form distinct pad; a greatly enlarged pair of apical subdigital lamellae, followed by a series of enlarged transverse lamellae, divided distally, single proximally; digits lying flat on substrate when viewed laterally; all digits clawed, the claws small, retractile and lying between the enlarged apical lamellae. Preanal pores present.” (cited after COUPER et al. 2007).

Definition (genus): A genus of the Diplodactylidae (sensu Han et al., 2004) distinguished from all related genera by the possession of enlarged juxtaposed dorsal scales approximately the same size as the ventrals (versus much smaller in related genera). Further dis- tinguished from other taxa formerly placed in Oedura (see above and below) by the combination of (1) moderate to large size (60–110 + mm), (2) karyotypic complement of 2n = 38, (3), posses- sion of one or more cloacal spurs (=cloacal spurs), and (4) dorsal pattern generally including a weak to bold series of transverse bands or disjunct blotches with no evidence of a vertebral stripe [OLIVER et al. 2012].

Diagnosis (genus): A genus of the Diplodactylidae distinguished from all related genera by the possession of enlarged juxtaposed dorsal scales approximately the same size as the ventrals (versus much smaller in related genera). Further distinguished from other taxa formerly placed in Oedura (Amalosia, Hesperoedura and Nebulifera; Oliver et al. 2012), by the combination of large size (adult SVL >60 mm), usually more than one cloacal spur per side, and dorsal pattern generally including a weak to bold series of transverse bands or disjunct blotches and no evidence of a vertebral stripe (OLIVER & DOUGHTY 2016).

Distribution: note that the distribution of O. mormorata has been restricted to the Northern Territory after splitting off other populations into different species (O. cincta, O. fimbria).

Karyotype: 2n = 38 (King 1987) 
References
  • Bedford, G.S.; Christian, K.A. 1998. Changes in Tail Color after Autotomy in the Gekkonid Lizard Oedura marmorata, from Australia. Jour. Int. Gecko Soc. 3 (3): 127-130
  • Boulenger, G.A. 1885. Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Bustard, H. R. 1970. Oedura marmorata a complex of Australian geckos (Reptilia: Gekkonidae). Senckenbergiana biologica 51: 21-40
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp.
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Cope, E.D. 1869. Observations on Reptiles of the old world. Proc. Acad. nat. Sci. Philadelphia 1868: 316-323 - get paper here
  • de Vis, C. W. 1884. Notes on the fauna of the Gulf of Carpentaria. Proc. Roy. Soc. Queensland 1: 154—160. - get paper here
  • De Vis, C. W. 1888. A contribution to the herpetology of Queensland. Proc. Linn. Soc. N. S. W. (2) 2: 811-826 [1887] - get paper here
  • Ford, J. 1963. The Reptilian Fauna of the Islands between Dongara and Lancelin, Western Australia. Western Australian Naturalist 8 (6): 135-142
  • Gray, J. E. 1842. Description of some hitherto unrecorded species of Australian reptiles and batrachians. Zoological Miscellany 2: 51—57 (London: Treuttel, Würtz & Co) - get paper here
  • Hagey TJ, Harte S, Vickers M, Harmon LJ, Schwarzkopf L 2017. There’s more than one way to climb a tree: Limb length and microhabitat use in lizards with toe pads. PLoS ONE 12(9): e0184641 - get paper here
  • Laube, A. & Langner, C. 2007. Die “Geckos” Australiens. Draco 8 (29): 4-21 - get paper here
  • Laube, A. & Langner, C. 2007. Australische Samtgeckos. Die Gattung Oedura. Natur und Tier Verlag (Münster), 64 pp. - get paper here
  • Longman, H. A. 1915. Reptiles from Queensland and the Northern Territory. Memoirs of the Queensland Museum 3: 30-34.
  • Oliver PM, McDonald PJ. 2016. Young relicts and old relicts: a novel palaeoendemic vertebrate from the Australian Central Uplands. R. Soc. open sci. 3: 160018 - get paper here
  • Oliver, Paul M.; Aaron M. Bauer, Eli Greenbaum, Todd Jackman, Tara Hobbie 2012. Molecular phylogenetics of the arboreal Australian gecko genus OeduraGray 1842 (Gekkota: Diplodactylidae): Another plesiomorphic grade?. Molecular Phylogenetics and Evolution 63 (2): 255-264 - get paper here
  • OLIVER, PAUL M.; PAUL DOUGHTY 2016. Systematic revision of the marbled velvet geckos (Oedura marmorata species complex, Diplodactylidae) from the Australian arid and semi-arid zones Zootaxa 4088 (2): 151–176 - get paper here
  • Rosauer, D.F., et al., 2016. Phylogeography, hotspots and conservation priorities: An example from the Top End of Australia. Biological Conservation, doi:10.1016/j.biocon.2016.05.002 - get paper here
  • Rösler, H. 2000. Studien an den Begattungsorganen der Geckos (Reptilia: Gekkota) - 3. Die Hemipenismorphologie von Arten der Gattungen Hoplodactylus FITZINGER 1843, Naultinus GRAY 1842, Oedura GRAY 1842, Rhacodactylus FITZINGER 1843 und Strophurus FITZINGER 1843 (Gek Gekkota 2: 220-248
  • Schmida, G. 2007. Betrachtungen zu den australischen Samtgeckos der Gattung Oedura. Draco 8 (29): 22-30 - get paper here
  • Wells, R. W. and Wellington, C. R. 1985. A classification of the Amphibia and Reptilia of Australia. Australian Journal of Herpetology, Supplementary Series, (1):1-61. - get paper here
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
 
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