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Oligodon nagao DAVID, NGUYEN, NGUYEN, JIANG, CHEN, TEYNIÉ & ZIEGLER, 2012

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Higher TaxaColubridae, Colubrinae, Colubroidea, Serpentes (snakes) 
Subspecies 
Common Names 
SynonymOligodon nagao DAVID, NGUYEN, NGUYEN, JIANG, CHEN, TEYNIÉ & ZIEGLER 2012 
DistributionN Vietnam (Cao Bang), Laos (Khammouane), SW China (Guangxi)

Type locality: Huu Lien forest, Huu Lung District, Lang Son Province, Vietnam, at elevation of about 300 m elevation. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
TypesHolotype. VNMN A.2012.1, adult male; collected by Tao Thien Nguyen on 30 June 2009.
Paratypes. Four specimens, all adult males: MNHN 2012.0216, from Huu Lien forest, Huu Lung District, Lang Son Province, Vietnam, at elevation of about 300 m a.s.l.; collected by Tao Thien Nguyen, 2 July 2009.—IEBR A.2012.6, from Duc Quang Commune, Ha Lang District, Cao Bang Province, Vietnam; collected by Truong Quang Nguyen et al., 14 October 2011 (22o42.859’N, 106o39.853’E, elevation 476 m).—KIZ 014591, from Nonggang National Nature Reserve, Longzhou County, Guangxi Autonomous Region, People’s Republic of China; collected by Tianbo Chen, October, 2011.—ZFMK 93281, from Ban Nathan, Hin Boun District, Khammouane Province, Laos PDR (17o58.854’N, 104o49.517’E, elevation 172 m); collected by Alexandre Teynié et al., 17 May 2012, during the mission “Opération Canopée, Inventaire de la Biodiversité Forestière du Laos 2012-2015”. 
CommentDiagnosis. A large species (TL up to at least 786 mm) of the genus Oligodon characterized by the combination of (1) 9 or 10 maxillary teeth, the last three or four strongly enlarged, (2) hemipenes not forked but divided into two lobes, thick and bulbous, reaching in situ the 16th SC, smooth, each lobe with a papilla, (3) 17–17–15 (or 17–15–15 DSR in one specimen); (4) cloacal plate entire, (5) complete complement of head scales, including 1 loreal on each side, (6) 8 supralabials (7 in one specimen), fourth and fifth (third and fourth in one specimen) entering orbit, (7) 184–193 ventrals in five males (females unknown), (8) tail relatively short in males (ratio TaL/TL: 0.135–0.146), (9) dorsal pattern made of a dark background colour with 27–37 darker, pale centered, butterfly-shaped blotches on the body and 5–8 on the tail, and (10) venter cream, heavily marked with dark pigmentation. 
References
  • DAVID, PATRICK; TRUONG QUANG NGUYEN, TAO THIEN NGUYEN, KE JIANG, TIANBO CHEN, ALEXANDRE TEYNIÉ & THO 2012. A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from northern Vietnam, southern China and central Laos. Zootaxa 3498: 45–62 - get paper here
  • VASSILIEVA, ANNA B.; PETER GEISSLER, EDUARD A. GALOYAN, NIKOLAY Jr A. POYARKOV, ROBERT WAYNE VAN DEV 2013. A new species of Kukri Snake (Oligodon Fitzinger, 1826; Squamata: Colubridae) from the Cat Tien National Park, southern Vietnam. Zootaxa 3702 (3): 233–246 - get paper here
 
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