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Ophisops elegans MÉNÉTRIES, 1832

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Higher TaxaLacertidae, Eremiadinae, Sauria, Lacertoidea, Squamata (lizards)
SubspeciesOphisops elegans basoglui BARAN & BUDAK 1978
Ophisops elegans blanfordi SCHMIDT 1939
Ophisops elegans budakibarani TOK, AFSAR, YAKIN, AYAZ & ÇİÇEK 2017
Ophisops elegans centralanatoliae BODENHEIMER 1944
Ophisops elegans ehrenbergii (WIEGMANN 1835)
Ophisops elegans elegans MÉNÉTRIÉS 1832
Ophisops elegans macrodactylus BERTHOLD 1840
Ophisops elegans schlueteri BOETTGER 1880 
Common NamesE: Snake-eyed Lizard
G: (Europäische) Schlangenauge(neidechse)
Russian: Стройная змееголовка 
SynonymOphisops elegans MÉNÉTRIES 1832: 63
Amystes ehrenbergi WIEGMANN 1835
Ophisops elegans — DUMÉRIL & BIBRON 1839: 259
Ophiops [sic] elegans — BERTHOLD 1840
Gymnops meizolepis STOLICZKA 1872 (fide SMITH 1935)
Ophisops elegans — ANDERSON 1872
Ophiops elegans — BOETTGER 1880
Ophiops elegans var. persicus BOULENGER 1918: 160
Ophiops elegans var. mizolepis — BOULENGER 1921 (fide SMITH 1935)
Ophisops elegans elegans — SMITH 1935: 379
Ophisops blanfordi SCHMIDT 1939
Ohisops [sic] elegans KHALAF 1961
Ophisops elegans — ENGELMANN et al 1993
Ophisops elegans — SCHLEICH, KÄSTLE & KABISCH 1996: 430
Ophisops elegans — ŠMÍD et al. 2014
Ophisops elegans — BAR et al. 2021

Ophisops elegans basoglui BARAN & BUDAK 1978
Ophisops elegans basoglui — SINDACO & JEREMČENKO 2008
Ophisops elegans basoglui — KUCHARZEWSKI 2016

Ophisops elegans blanfordi SCHMIDT 1939
Ophisops blanfordi SCHMIDT 1939
Ophisops blanfordi — HELLMICH 1959
Ophisops elegans blanfordi — SINDACO & JEREMČENKO 2008
Ophisops blanfordi — MOHAMMED et al. 2017

Ophisops elegans budakibarani TOK, AFSAR, YAKIN, AYAZ & ÇİÇEK 2017

Ophisops elegans centralanatoliae BODENHEIMER 1944: 33
Ophisops elegans centralanatoliae — SINDACO & JEREMČENKO 2008
Ophisops elegans centralanatoliae — TAYHAN et al. 2016

Ophisops elegans ehrenbergii (WIEGMANN 1835)
Amystes Ehrenbergii WIEGMANN 1835
Ophisops elegans ehrenbergi — WERNER 1935: 106
Ophisops elegans ehrenbergii — BIRD 1936
Ophisops elegans erhenbergii [sic] — ANGEL 1936: 111
Ophisops elegans ehrenbergii — SCHMIDT 1939
Ophisops elegans ehrenbergii — BISCHOFF 1991
Ophisops elegans ehrenbergii — SINDACO & JEREMČENKO 2008
Ophisops elegans cf. ehrenbergii — SINDACO et al. 2014
Ophisops elegans ehrenbergii — MOHAMMED et al. 2017

Ophisops elegans macrodactylus BERTHOLD 1840
Ophisops macrodactylus BERTHOLD 1840: 58
Ophisops macrodactylus — BERTHOLD 1846: 15
Ophisops elegans macrodactylus — ÇEVIK 1999
Ophisops elegans macrodactylus — SCHLÜTER 2004
Ophisops elegans macrodactylus — SINDACO & JEREMČENKO 2008
Ophisops elegans macrodactylus — SCHLÜTER 2010
Ophisops elegans macrodactylus — DEICHSEL 2013
Ophisops macrodactylus — BÖHME 2014: 151
Ophisops elegans macrodactylus — KWET & TRAPP 2014
Ophisops elegans macrodactylus — BAMANN et al. 2021

Ophisops elegans schlueteri BOETTGER 1880
Ophiops [sic] elegans schlueteri BOETTGER 1880: 176
Ophisops elegans schlueteri — SCHMIDT 1939
Ophisops elegans schlueteri — ESTERBAUER 1985
Ophisops elegans schlueteri — SINDACO & JEREMČENKO 2008
Ophisops elegans schlueteri — SCHNEIDER 2020 
DistributionE Republic of Georgia, Armenia, Azerbaijan,
SE Bulgaria, NE Greece (Lesbos, Limnos, Chios, Samos, Samothraki, Agathonisi, Psara, Kalymnos), Cyprus, Turkey,
Algeria, Egypt, W Syria, Lebanon, Israel, W Jordan,
Iraq, Iran (Kavir desert), Russia (Chechnya and Dagestan)

basoglui: Turkey (S Anatolia, Alanya area)

blanfordi: Mesopotamia, SE Iraq, SW Iran.

centralanatoliae: C/E/S Turkey (C Anatolia)

ehrenbergi: Levant, Greece (Kalymnos, Lesbos etc.), S Turkey (incl Anatolia); N Syria, Iraq

elegans: Turkey (incl Anatolia); Russia (Chechnya and Dagestan), N Iraq, N/C Iran, N Pakistan; Type locality: Baku, Azerbaijan

macrodactylus: W Turkey (Anatolia), Greece (Lesbos, Samos, Rhodes etc.); Type locality: Constantinopel [= Istanbul]

schlueteri: Cyprus  
TypesSyntype: USNM 21396
Holotype: ZDEU (also given as SZE) 167/1975.1, Turkey, Alanya [basoglui]
Holotype: FMNH 19721, paratypes: MSNG [blanfordi]
Holotype: COMU-ZDEU 2017/10 (COMU = Çanakkale Onsekiz Mart University, which is different from ZDEU) [budakibarani]
Holotype: ZMB [ehrenbergii]
Holotype: ZFMK 21219, cited erroneously as ZDFMK by Böhme 2010 [macrodactylus]
Syntypes: ZMUI ("Zoological Institutes at Istanbul University and Ankara (Y.Z.E.)", BMNH (123+ specimens) [centralanatoliae]
Syntypes: BMNH [persicus]
Lectotype: SMF 13841 [schlueteri] 
DiagnosisDiagnosis (genus): Lower eyelid fused with upper, with large transparent disc (Fig. 123); ventral plates smooth; collar weakly defined, or absent in middle; subdigital lamellae keeled; dorsal scales rhombic, imbricate, strongly keeled; femoral pores present (Anderson 1999: 254).

Diagnosis: Upper head shields smooth, 27-28 scales and plates round body; snout shorter than breadth o f head across eyes; scales on nape small and granular; supraoculars sepa- rated from superciliaries by a series of small granules. (Anderson 1999: 255)

Color pattern: Olive-greenish or brownish above, with 2 light dorsolateral stripes, upper running from supraciliary margin to tail; lower running from below eye through ear, along flank to hind limb; upper margin ofdorsolateral stripe spotted with black; light stripes occasionally absent in adults; black vertebral streak often present, at least on neck; a series of small vertebral spots occasionally present; supralabials, sides of neck, and interval between light stripes spotted with black; venter greenish white; distal half of tail often reddish. Hatchlings with contrasting chocolate brown and bright yellow stripes. (Anderson 1999: 255)

Size: Snout-vent length to 62 mm. (Anderson 1999: 255)

Diagnosis (budakibarani): This new subspecies can be distinguished from Ophisops elegans basoglui known from southern Anatolia and O. e. macrodactylus (in both sexes) mainly distributed in the western Anatolia by absence of coloration on the venter during the breeding season. Ophisops elegans budakibarani is differentiated from nominate subspecies and O. e. centralana- tolia in the north of the its distribution by the presence of dark brown or less distinct reddish brown temporal band in- stead of very distinct reddish brown one. To take into the consideration the longitudinal row of dorsal scales + ventral plates at mid-trunk is the main diagnostic character among subspecies, O. e. budakibarani is easily distinguishable from the O. e. basoglui by having a higher value, while it has lower value than O. e. centralanatoliae.

Subspecies are differentiated by the following characters, according to Darevsky & Beutler 1981:

O. e. elegans: Jederseits zwei Postnasalia, dunkler Vertebralstreif meist vorhandenfum 33.(28-40) Schuppen um die Körpermitte, 9-13 Femoral­ poren, um 40 (26-60) Temporalschuppen; Kopfseiten- und Kehlfärbung weiß.

O. e. ehrenbergii: Jederseits zwei Postnasalia, Vertebralstreifen meist fehlend, 24-36 Schuppen um die Rumpfmitte, 9-13 Femoralporen, 17-47 Schuppen in der Temporalregion. Kopfseite und Kehle lebender Männchen gelb.

O. e. basoglui: Voriger Form ähnlich, Vertebralstreif fehlt ebenfalls meist, aber beim lebenden 3 ist die ganze Unterseite intensiv gelb, beim Weibchen die Kehle.

O. e. centralanatoliae: Jederseits zwei Postnasalia, Vertebralstreif oft vorhanden, um 38 (34-43) Schuppen um die Körpermitte, 9-12 Femoral­ poren, um 48 (29-70) Temporalschuppen.

O. e. schlueteri: Jederseits zwei Postnasalia, Vertebralstreif gewöhnlich fehlend, um 43 (38-49) Schuppen um die Körpermitte, 12-14 Femoralporen, um 75 (50-104) Temporalschuppen.

O. e. blanfordi: Jederseits ein Postnasale, V ertebralstreif meist vorhanden, 29-38 Schuppen um die Körpermitte. 
CommentSynonymy: O. e. persicus was considered a synonym of O. elegans elegans, e.g. by Anderson 1999: 254, who rejects subspecies in O. elegans in general.

Subspecies: the current subspecies classification of O. elegans does not match the molecular clades found by Montgelard et al. 2021 and may need to be revised.

Distribution: Not listed for India by DAS 1997. Not in Libya fide Bauer et al. 2017. Distribution: See map in SMID et al. 2014 for distribution in Iran. See map in SMID et al. 2014 for distribution in Iran. See for maps of subspecies. For a map of type localities of subspecies see Montgelard et al. 2021: Fig. 1b. See Bar et al. 2021 for a map.

Type species: Ophisops elegans MÉNÉTRIES 1832: 63 is the type species of the genus Ophisops MÉNÉTRIES 1832.

Comparisons: Patel & Vyas 2020 presented a comparison of characters in Indian Ophisops. 
EtymologyThe name elegans (Latin) means tasteful, choice, fine, select or elegant and appears to refer to the color pattern or body shape. 
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