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Higher TaxaViperidae, Crotalinae, Colubroidea, Alethinophidia, Serpentes, Squamata (snakes) 
Common NamesEmerald Horned Pitviper 
SynonymOphryacus smaragdinus GRÜNWALD, JONES, FRANZ-CHÁVEZ & AHUMADA-CARILLO 2015
Ophryacus undulatus — FLORES-VILLELA et al. 1992: 249.
Ophryacus undulatus — CAMARILLO 1995: 196.
Ophryacus undulatus — CAMPBELL & LAMAR 2004: 454 (in part).
Ophryacus undulatus — MENDOZA-PAZ et al. 2006: 109 (in part).
Ophryacus undulatus — RAMÍREZ-BAUTISTA et al. 2010: 77
Ophryacus undulatus — SOLANO-ZAVALETA et al. 2010: 381
Ophryacus undulatus — RAMÍREZ-BAUTISTA et al. 2014: 177 
DistributionMexico (Veracruz, Hidalgo, Puebla, Oaxaca), elevation 1,400 - 2,400 m

Type locality: Los Ocotes (20.601542, -98.469708; datum WGS 84; elev. 2,150 m asl), Zilacatipan, Municipio de Huayacocotola, Veracruz, Sierra Madre Oriental, Mexico  
TypesHolotype: MZFC 29290, adult female, collected by Jason M. Jones and Iván T. Ahumada-Carrillo on 2 November 2013.
Paratypes: Seven specimens. MEXICO: HIDALGO: El Potrero, Municipio de Tenango de Doria (20.314922, -98.229653; datum WGS 84; elev. 2,200 m), an adult male collected 25 August 1990 by students of the Escuela Nacional Estatal y Profesional de Iztacala, UNAM (MZFC 21140); “Doria,” Municipio de Tengano de Doria (20.314922, -98.229653; datum WGS 84; elev. 2,200 m), an adult male collected on 14 July 1990 by Efraín Hernández-García (MZFC 04825). OAXACA: San Martín Caballero, Municipio de San José Tenango (18.108175, -96.633981; datum WGS 84; elev. 1,360 m), an adult male collected 25 May 1994 by Mario Mancilla-Moreno (MZFC 13556); San Martín Caballero, Municipio de San José Tenango (18.108175, -96.633981; datum WGS 84; elev. 1,360 m), a juvenile female collected 28 May 1994, collector unknown (MZFC 13557). PUEBLA: Xucayucan, Municipio de Tlatlauquitepec (19.895744, -97.471700; datum WGS 84; elev. 1,630 m), a juvenile female collected 27 May 2005 by Israel Solano-Zavaleta (MZFC 17663). VERACRUZ: Cerro La Campana, Ticuautipan, Municipio de Xico (19.439500, -97.078078; datum WGS 84; elev. 2,045 m), a subadult female collected on 26 October 2003 by Itzel Durán Fuentes (MZFC 20582); and Zilacatipan, Municipio de Huayacocotla (20.655761, -98.424894; datum WGS 84; elev. 1,790 m), a subadult female, collected on 2 November 2013 by Jason M. Jones and Iván T. Ahumada-Carrillo (MZFC 29289). 
EtymologyThe specific epithet is derived from the Latin word smaragdinus, an adjective meaning “emerald-green,” in reference to the emerald green ground coloration exhibited by most individuals of this species. 
  • Camarillo,R.J.L. 1995. Distribution records for some amphibians and reptiles from Mexico. Bull. Maryland Herp. Soc. 31 (4): 195-197
  • Campbell, Jonathan A. and William W. Lamar 2004. The Venomous Reptiles of the Western Hemisphere, 2 vols. Comstock (Cornell University Press), Ithaca, NY, 962 pp. [review in Science 305: 182]
  • Flores-Villela, O., F. Mendoza, E. Hernández, M. Mancilla, E. Godínez, and I. Goyenechea-Mayer. 1992. Ophryacus undulatus in the Mexican state of Hidalgo. Texas Journal of Science 44: 249–250
  • Grünwald, Christoph I.; Jason M. Jones, Hector Franz-Chávez, and Iván T. Ahumada-Carrillo 2015. A new species of Ophryacus (Serpentes: Viperidae: Crotalinae) from eastern Mexico, with commentson the taxonomy of related pitvipers. Mesoamerican Herpetology 2 (4): 388 - get paper here
  • Mendoza-Paz, R. F., F. Mendoza-Quijano, R. W. Bryson, and E. N. Smith. 2006. Geographical Distribution. Ophryacus undulatus (Mexican Horned Pitviper). Herpetological Review 37: 109 - get paper here
  • Ramírez-Bautista, A., U. Hernández-Salinas, F. Mendoza-Quijano, R. Cruz-Elizalde, B. P. Stephenson, V. D. Vite-Silvaand A. Leyte-Manriquez. 2010. Lista Anotada de los Anfibios y Reptiles del Estado de Hidalgo, México. Universidad Autónoma del Estado de Hidalgo, Pachuca, Hidalgo, and Comisión Nacional para el Conocimiento y Uso de la Biodiversidad, México D.F., Mexico
  • Ramírez-Bautista, Aurelio, Uriel Hernández-Salinas, <br />Raciel Cruz-Elisade, Christian Berriozabal-Islas, Daniel Lara-Tufiño, Irene Goyenechea Mayer-Goenechea, and Jesús M. Castillo-Cerón 2015. Los Anfibios y Reptiles de Hidalgo, México: Diversidad, Biogeografía y Conservación. Eagle Mountain Publishing, xiii + 387 pp.
  • Solano-Zavaleta, Israel, Andrés Alberto Mendoza-Hernández and Luis Canseco-Márquez. 2010. Geographic distribution: Ophryacus undulatus. Herpetological Review 41 (3): 381 - get paper here
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