Ophryacus undulatus (JAN, 1859)

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Higher Taxa	Viperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies
Common Names	E: Slender-horned Pitviper, Mexican Horned Pit Viper S: Cornuda Mexicana
Synonym	Trigonocephalus (Atropos) undulatus JAN 1859 Teleuraspis undulatus — GARMAN 1884: 126 Ophryacus undulatus — FERRARI-PÉREZ 1886 (nom. nud.) Lachesis undulatus BOULENGER 1896 Trimeresurus undulatus — SMITH 1941: 63 Ophryacus undulatus — CAMPBELL & LAMAR 1989: 311 Ophryacus undulatus — LINER 1994 Ophryacus undulatus — WELCH 1994: 95 Ophryacus undulatus — MCDIARMID, CAMPBELL & TOURÉ 1999: 314 Ophryacus undulatus — JADIN et al. 2011 Ophryacus undulatus — WALLACH et al. 2014: 508
Distribution	Mexico (Guerrero, Oaxaca: Sierra Madre Oriental, Mesa del Sur, Sierra Madre del Sur; Puebla, Hidalgo, Veracruz) Type locality: "Messico" [Mexico]. Proposed restriction to "Orizaba, Veracruz, Mexico" by Smith and Taylor, 1950: 350.
Reproduction	ovovivparous
Types	Syntypes: lost, formerly MSNM (2), Milan Museum, lost during WW II (S. Scali, pers. comm., cited in GRÜNWALD et al. 2015).
Diagnosis	Additional details, e.g. a detailed description or comparisons (3067 characters), are available for collaborators and contributors. Please contact us for details.
Comment	Venomous! Habitat: forests Synonyms: Bothrops sphenophrys SMITH 1960 has been removed from the synonymy of O. undulatus by GRÜNWALD et al. 2015. Sympatry: O. sphenophrys and O. undulatus have been collected within 7 km of one another. Type species: Trigonocephalus undulatus JAN 1859 is the type species of the genus Ophryacus COPE 1887.
Etymology	Named after Latin undulatus, that has a wave-like form of the dorsal pattern. The genus name is derived from the Greek ophrys, meaning brow, and the Latin acus, meaning pointed, obviously in reference to the distinctive supraocular spine-like scale.
References	Camarillo,R.J.L. 1995. Distribution records for some amphibians and reptiles from Mexico. Bull. Maryland Herp. Soc. 31 (4): 195-197 - get paper here Campbell, J.A. & Lamar, W.W. 1989. The Venomous Reptiles of Latin America. Comstock Publishing/Cornell University Press, Ithaca Canseco-Márquez, L., & Gutiérrrez-Mayén, M.G. 2010. Anfibios y reptiles del Valle de Tehuacán-Cuicatlán. Comisión Nacional para el conocimiento y uso de la biodiversidad, México D.F., Mexico, 302 pp - get paper here Canseco-Marquez,L.; Gutierrez-Mayen,G. & Salazar-Arenas,J. 2000. New records and range extensions for amphibians and reptiles from Puebla, México. Herpetological Review 31 (4): 259-263 - get paper here Casas-Andreu, G., F.R. Méndez-De la Cruz and X. Aguilar-Miguel. 2004. Anfibios y Reptiles; pp. 375–390, in A.J.M. García-Mendoza, J. Ordoñez and M. Briones-Salas (ed.). Biodiversidad de Oaxaca. Instituto de Biología, UNAM-Fondo Oaxaqueño para la Conservación de la Naturaleza-World Wildlife Fund, México, D. F. Garman, Samuel 1884. The reptiles and batrachians of North America. Mem. Mus. comp. Zool, Cambridge (Massachusetts), 8 (3): xxxiv + 185 pp. [1883] [CNAH reprint 10] - get paper here Grünwald, Christoph Imre, Nadia Pérez-Rivera, Iván Trinidad Ahumada-Carrillo, Héctor Franz-Chávez and Brandon Thomas La Forest. 2016. New distributional records for the herpetofauna of Mexico. Herpetological Review 47 (1): 85-90 - get paper here Grünwald, Christoph I.; Jason M. Jones, Hector Franz-Chávez, and Iván T. Ahumada-Carrillo 2015. A new species of Ophryacus (Serpentes: Viperidae: Crotalinae) from eastern Mexico, with commentson the taxonomy of related pitvipers. Mesoamerican Herpetology 2 (4): 388 - get paper here Heimes, P. 2016. Snakes of Mexico. Chimaira, Frankfurt, 572 pp JADIN, ROBERT C.; ERIC N. SMITH and JONATHAN A. CAMPBELL 2011. Unravelling a tangle of Mexican serpents: a systematic revision of highland pitvipers. Zoological Journal of the Linnean Society 163: 943–958 - get paper here Jan, G. 1859. Plan d'une iconographie descriptive des ophidiens et description sommaire de nouvelles espèces de serpents. Rev. Mag. Zool., Paris (2) (11-12): 122-130 - get paper here Lemos-Espinal, Julio A., Geoffrey R. Smith 2015. Amphibians and reptiles of the state of Hidalgo, Mexico. Check List 11 (3): 1642 - get paper here Mata-Silva, Vicente, Jerry D. Johnson, Larry David Wilson and Elí García-Padilla. 2015. The herpetofauna of Oaxaca, Mexico: composition, physiographic distribution, and conservation status. Mesoamerican Herpetology 2 (1): 6–62 - get paper here McDiarmid, R.W.; Campbell, J.A. & Touré,T.A. 1999. Snake species of the world. Vol. 1. [type catalogue] Herpetologists’ League, 511 pp. O’Shea, M. 2018. The Book of Snakes. Ivy Press / Quarto Publishing, London, - get paper here Palacios-Aguilar, Ricardo & OSCAR FLORES-VILLELA 2018. An updated checklist of the herpetofauna from Guerrero, Mexico. Zootaxa 4422 (1): 1-24 - get paper here Parkinson,C.L. 1999. Molecular systematics and biogeographical history of pitvipers as determined by mitochondrial ribosomal DNA sequences. Copeia 1999 (3): 576-586 - get paper here Ramírez-Jiménez, Francisco, Pablo R. Simón-Salvador, Elí García-Padilla, Vicente Mata-Silva and Larry David Wilson. 2016. Ophryacus undulatus (Jan, 1859). Mexico, Oaxaca. Mesoamerican Herpetology 3(4): 1069 - get paper here Smith, H.M. 1941. Notes on Mexican snakes of the genus Trimeresurus. Zoologica 26: 61-64. - get paper here Solano-Zavaleta, Israel, Andrés Alberto Mendoza-Hernández and Luis Canseco-Márquez. 2010. Geographic distribution: Ophryacus undulatus. Herpetological Review 41 (3): 381 - get paper here Taggart, Travis W., Brian I. Crother, and Mary E. White 2001. Palm-pitviper (Bothriechis) phylogeny, mtDNA, and consilience. Cladistics 17: 355-370 - get paper here Torres-Hernández, LA, Ramírez-Bautista A, Cruz-Elizalde R, Hernández-Salinas U, Berriozabal-Islas C, DeSantis DL, Johnson JD, Rocha A, García-Padilla E, Mata-Silva V, Fucsko LA, and Wilson LD. 2021. The herpetofauna of Veracruz, Mexico: composition, distribution, and conservation status. Amphibian & Reptile Conservation 15(2) [General Section]: 72–155 - get paper here Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp. Woolrich-Piña, G. A., E. García-Padilla, D. L. DeSantis, J. D. Johnson, V. Mata-Silva, and L. D. Wilson 2017. The herpetofauna of Puebla, Mexico: composition, distribution, and conservation status. Mesoamerican Herpetology 4(4): 791–884 - get paper here
External links	IUCN Red List National Center for Biotechnology Information Google images

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