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Pantherophis guttatus (LINNAEUS, 1766)

IUCN Red List - Pantherophis guttatus - Least Concern, LC

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Higher TaxaColubridae, Colubrinae, Lampropeltini, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Red Cornsnake, Eastern Corn snake
G: Gewöhnliche Kornnatter 
SynonymColuber guttatus LINNAEUS 1766: 385
Coluber maculatus BONNATERRE 1790: 19
Coluber compressus DONNDORF 1798: 206
Coluber carolinianus SHAW 1802: 460
Coluber molossus DAUDIN 1803: 269
Coluber pantherinus DAUDIN 1803: 318
Coluber floridanus HARLAN 1827: 360
Pantherophis guttata FITZINGER 1843
Scotophis guttatus — BAIRD & GIRARD 1853: 78
Elaphis guttatus — DUMÉRIL, BIBRON & DUMÉRIL 1854: 273
Coryphodon Pantherinus DUMÉRIL, BIBRON & DUMÉRIL 1854: 181
Elaphis guttatus — GARMAN 1884: 55
Coluber guttatus sellatus COPE 1888: 387
Coluber rosaceus COPE 1888: 388
Coluber guttatus — BOULENGER 1894: 39
Callopeltis guttatus — LOENNBERG 1894
Coluber laetus — BOULENGER 1894: 49
Coluber guttatus — COPE 1900: 833
Coluber guttatus sellatus — COPE 1900: 836
Coluber rosaceus — COPE 1900: 837
Elaphe laeta — BURT 1935
Elaphe laeta — WOODBURY 1942
Elaphe emoryi intermontana WOODBURY & WOODBURY 1942
Elaphe laeta laeta — TAYLOR 1949: 203
Elaphe guttata — STEBBINS 1985: 186
Elaphe guttata guttata — CONANT & COLLINS 1991: 194
Elaphe guttata — SCHWARTZ & HENDERSON 1991: 600
Elaphe guttata — LINER 1994
Elaphe guttata — SCHULZ 1996: 281
Elaphe guttata rosacea — COWLES 1996
Elaphe guttata guttata — TENNANT & BARTLETT 2000: 392
Pantherophis guttatus — UTIGER et al. 2002
Elaphe guttata — GIBBONS & DORCAS 2005
Pituophis guttatus — BURBRINK 2007
Pantherophis guttatus — COLLINS & TAGGART 2008
Pantherophis guttatus — WALLACH et al. 2014: 529
Pantherophis gutattus — KRAVCHUK & WATSON 2020 (in error) 
DistributionUSA (Arkansas, Mississippi, Alabama, South Carolina, North Carolina, Kentucky, Tennessee, Nebraska, Virginia, Maryland, Delaware, West Virginia, New Jersey, Florida, Texas, Louisiana, Georgia),
Cayman I, US Virgin Islands, Anguilla, Antigua, St. Bartélmy (Saint Barthélemy) (POWELL & HENDERSON 2003)

Introduced to Brazil (Bahia)

Type locality: “Carolina” (Linnaeus, 1766); restricted to USA: South Carolina, Charleston County, vicinity of Charleston (Dowling, 1952)  
Reproductionoviparous 
TypesHolotype: UUZM (= ZMUU) 147 (Zoological Museum of Uppsala)
Holotype: ANSP 3875; east Florida [Coluber floridanus HARLAN 1827]
Syntypes: USNM 9692, 5507 [Coluber guttatus sellatus]
Holotype: USNM 14418 [Coluber rosaceus]
Syntype: USNM 391 [Coryphodon pantherinus] 
DiagnosisAdditional details, e.g. a detailed description or comparisons (909 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSubspecies: Elaphe guttata emoryi (Baird & Girard 1853) has been elvated to full species status.

Type species: Coluber guttatus LINNAEUS 1766 is the type species of the genus Pantherophis FITZINGER 1843: 25.

Hybrids: BRÖER (1977) reports a hybrid between E. guttata guttata and E. obsoleta quadrivittata. KREUTZ (2005) reports hybrids between E. guttata and Lampropeltis getulus californiae, between E. guttata and L. pyromelana, and between E. guttata and L. triangulum sinaloae. Bechtel & Mountain (1960) report hybrids between Elaphe guttata and E. obsoleta rossaleni.

Genome: a draft genome of this species has been published by Ullate-Agote et al. 2015.

Synonymy partly after BURBRINK 2002.

Illustration: SCHMDIT (2004) illustrates a striped mutation.

Phylogenetics: see Dahn et al. 2018 for a phylogeny of Lampropeltini, but also see Burbrink et al. 2020, Hillis & Wüster (2021), and Burbrink et al. 2021 for the relationships among obsoletus, alleghaniensis, quadrivittatus, and bairdi.

Habitat: partly arboreal (Harrington et al. 2018). 
EtymologyNamed after Latin guttatus, a, um, ‘spotted, speckled’, in reference to the spotted coloration of this taxon.

The genus name was derived from the Greek words panther, meaning "panther" and ophis, meaning "snake," probably in reference to the spotted and colorful pattern of the type species. 
References
  • Abraham, G. 1980. Elaphe guttata guttata (KORNNATTER). Sauria 2 (3): 1-2 - get paper here
  • Allen, Morrow J. 1932. A survey of the Amphibians and reptiles of Harrison County, Mississippi. American Museum Novitates (542): 1-20 - get paper here
  • Anonymous 2007. Snakes of New Jersey. New Jersey Division of Fish and Wildlife
  • Augstenová, Barbora; Sofia Mazzoleni, Lukáš Kratochvíl Orcid and Michail Rovatsos 2017. Evolutionary Dynamics of the W Chromosome in Caenophidian Snakes. Genes 2018, 9(1), 5; doi:10.3390/genes9010005 - get paper here
  • Austin, James D.;Gregory, Patrick T. 1999. Relative roles of thermal and chemical cues in the investigative behavior of prey in colubrid (Elaphe guttata and Lampropeltis getulus) and Boid (Python regius) snakes. Herpetological Natural History 6 (1): 47-50 [1998]
  • Baird, S. F. and C. Girard. 1853. Catalogue of North American Reptiles in the Museum of the Smithsonian Institution. Part 1.-Serpents. Smithsonian Inst., Washington, xvi + 172 pp. - get paper here
  • Beane, J C 2019. New geographic distribution records for reptiles from North Carolina, USA. Herpetological Review 50 (3): 538-542 - get paper here
  • Bechtel, H. B.;Bechtel, E. 1989. Color mutations in the corn snake (Elaphe guttata guttata): Review and additional breeding data. Journal of Heredity 80: 272-276 - get paper here
  • Bechtel, H.B. & Mountain, J. 1960. Interspecific Hybridization between Two Snakes of the Genus Elaphe. Copeia 1960 (2): 151-153 - get paper here
  • Bird, William M., Philip Peak, and Danna L. Baxley. 2015. Natural hisotry of an allopatric population of Red Cornsnakes, Pantherophis guttatus (Linnaeus, 1766) in central Kentucky, USA. The Journal of North American Herpetology 2015(1):6-11 - get paper here
  • Bird, William M., Philip Peak, and Danna L. Baxley. 2015. Natural history of an allopatric population of Red Cornsnakes, Pantherophis guttatus (Linnaeus, 1766) in central Kentucky, USA. The Journal of North American Herpetology 2015(1):6-11 - get paper here
  • Blom, M. 2003. Algemene verzorging en kweek van Lampropeltis en Elaphe soorten. Lacerta 61 (1): 32-39 - get paper here
  • Boistel, Renaud;, Anthony Herrel; Renaud Lebrun, Gheylen Daghfous, Paul Tafforeau, Jonathan B. Lososô and Bieke Vanhooydonck|| 2011. Shake Rattle and Roll: The Bony Labyrinth and Aerial Descent in Squamates. Integrative and Comparative Biology, doi:10.1093/icb/icr034 - get paper here
  • Bonnaterre, P.J. 1790. Ophiologie, in Tableau encyclopédique et méthodique des trois règnes de la nature. Panconoke, Paris, XLIV + 76 pp + 43 plates - get paper here
  • Boulenger, George A. 1894. Catalogue of the snakes in the British Museum (Natural History). Volume II., Containing the Conclusion of the Colubridæ Aglyphæ. British Mus. (Nat. Hist.), London, xi, 382 pp. - get paper here
  • Brady, M. K. 1933. The third specimen of Elaphe rosacea (Cope). Proc. Biol. Soc. Washington 46: 153-154 - get paper here
  • Bröer, W. 1978. Bastarde bei zwei Elaphe-Arten. Salamandra 14 (2): 63-68 - get paper here
  • Burbrink, F.T. 2002. Phylogeographic analysis of the cornsnake (Elaphe guttata) complex as inferred from maximum likelihood and Bayesian analyses. Molecular Phylogenetics and Evolution 25 (3): 465-476 - get paper here
  • Burbrink, Frank T. and Robin Lawson 2007. How and when did Old World ratsnakes disperse into the New World? Molecular Phylogenetics and Evolution 43 (1): 173-189 - get paper here
  • Burt, Charles E. 1933. A collection of amphibians and reptiles from southern Missouri. American Midland Naturalist 14 (2): 170-173 - get paper here
  • Burt, Charles E. 1935. Further records of the ecology and distribution of amphibians and reptiles in the middle west. American Midland Naturalist 16 (3): 311-336 - get paper here
  • Camper, Jeffrey D. 2019. The Reptiles of South Carolina. University of South Carolina Press, 288 pp. [review in Copeia 107 (3): 590, 2019] - get paper here
  • Christman, S. P. 1980. Patterns of geographic variation in Florida snakes. Bulletin of the Florida State Museum of Biological Sciences 25 (3): 1 (157?)-256 - get paper here
  • Collins, Joseph T. & Travis W. Taggart 2008. An alternative classification of the New World Rat Snakes (genus Pantherophis [Reptilia: Squamata: Colubridae]). Journal of Kansas Herpetology (26): 16-18 - get paper here
  • Conant,R. & Collins,J.T. 1991. A Field Guide to Reptiles and Amphibians of Eastern/Central North America, 3rd ed. Houghton Mifflin (Boston/New York), xx + 450 p.
  • Cope, E.D. 1889. On the snakes of Florida. Proc. US Natl. Mus. 11: 381-394 [1888] - get paper here
  • Cope, E.D. 1900. The crocodilians, lizards and snakes of North America. Ann. Rep. U.S. Natl. Mus. 1898: 153-1270 - get paper here
  • Cowles, J.H. 1996. The case of the missing color: the "piebald" Rosy Rat Snake. Sonoran Herpetologist 9 (7):67. - get paper here
  • Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92
  • Dahn, H. A., Strickland, J. L., Osorio, A., Colston, T. J., & Parkinson, C. L. 2018. Hidden diversity within the depauperate genera of the snake tribe Lampropeltini (Serpentes, Colubridae). Molecular Phylogenetics and Evolution - get paper here
  • Daltry, Jennifer C. 2007. An introduction to the herpetofauna of Antigua, Barbuda and Redonda, with some conservation recommendations. Applied Herpetology 4 (2): 97-130. - get paper here
  • Daudin, F. M. 1803. Histoire Naturelle Generale et Particuliere des Reptiles. Vol. 6. F. Dufart, Paris, 447 pp. - get paper here
  • Degenhardt, William G.; C. W. Painter, and A. H. Price 1996. Amphibians and reptiles of New Mexico. Univ. New Mexico Press, 431 pp.
  • DeGregorio, Brett A.; Patrick J. Weatherhead, and Jinelle H. Sperry 2016. Ecology and Predation Behavior of Corn Snakes (Pantherophis guttatus) on Avian Nests. Herp. Cons. Biol. 11 (1) - get paper here
  • Donndorff, JOHANN AUGUST 1798. Zoologische Beyträge zur xiii. Ausgabe des Linnéischen Natursystems. Vol. 3. Leipzig, Weidmannschen, vi+980 pp. - get paper here
  • Douglas, D.A.; Janke, A. & Arnason, U. 2006. A mitogenomic study on the phylogenetic position of snakes. Zoologica Scripta 35: 545–558 - get paper here
  • Dowling, Herndon G. 1951. A taxonomic study of the ratsnakes, genus Elaphe Fitzinger. I. The status of the name Scotophis laetus Baird and Girard (1853). Copeia 1951 (1): 39-44 - get paper here
  • Drees, Stephan D 2014. Ein Blick auf die Farbzucht bei Pantherophis guttatus. Draco 16 (57): 60-69 - get paper here
  • Duméril, A.M.C., G. BIBRON & A.H.A. DUMÉRIL 1854. Erpétologie générale ou Histoire Naturelle complète des Reptiles. Vol. 7 (partie 1). Paris, xvi + 780 S. - get paper here
  • Dunaev E.A., Orlova V.F. 2003. Diversity of snakes (on the materials of the exposition of the Zoological Museum of the Moscow State University). Moscow: Moscow State University Press, 376 pp
  • Enge, Kevin M. 2009. Venomous and non-venomous snakes of Florida. Publication of the Florida Fish & Wildlife Conservation Commission. 16 pp.
  • Ernst, C.H., & Barbour, R.W. 1989. Snakes of eastern North America. George Mason Univ. Press, Fairfax, VA 282 pp.
  • Fonseca, E., R. Marques & M. S. Tinôco 2014. New records of Pantherophis guttatus (Squamata: Colubridae) in the state of Bahia, an alien species to Brazil. Salamandra 50 (4): 241-244 - get paper here
  • Garman, Samuel 1884. The reptiles and batrachians of North America. Mem. Mus. comp. Zool, Cambridge (Massachusetts), 8 (3): xxxiv + 185 pp. [1883] [CNAH reprint 10] - get paper here
  • Gerald, Gary W; Emma D Wass, 2019. Correcting for individual quality reveals trade-offs in performance among multiple modes of limbless locomotion in snakes. Biological Journal of the Linnean Society, Volume 128, Issue 2, October 2019, Pages 278–288, - get paper here
  • Gibbons, J.W. & Mike Dorcas 2005. Snakes of the Southeast. University of Georgia Press, 253 pp.
  • Glaß, M, & Bohle, D. 2012. Haltungsgrundlagen der Kornnatter. Vivaria-Verlag, Meckenheim, 160 pp.
  • Glaß, M, & Bohle, D. 2012. Farb- und Zeichnungsvarianten der Kornnatter. Vivaria-Verlag, Meckenheim, 64 pp.
  • Glaß, Michael 2009. Kornnattern. Vivaria Verlag, 144 pp.
  • GODLEY, J. S., J. L. HOFKES, AND C. M. REILLY 2020. Minimum Density Estimates of Terrestrial Snakes in the Munson Sandhills Adjoining Apalachicola National Forest, Florida. Herpetological Review 51: 221 - 226.
  • Green, N.B., & Pauley, T.K. 1987. Amphibians and reptiles in West Virginia. Univ. of Pittsburgh Press, Pittsburgh, 241 pp.
  • GRISNIK, M., AND R. HANSCOM 2020. New County Records for Reptiles and Amphibians from Middle Tennessee’s Cumberland Plateau. Herpetological Review 51: 282-284.
  • Grove, Barbara and David A. Steen. 2015. Geographic Distribution: Pantherophis guttatus (red cornsnake). Herpetological Review 46 (4): 575 - get paper here
  • Groves, J.D. 1967. Notes on Albinism in Elaphe guttata. Bull. Maryland Herp. Soc. 3 (2): 26 - get paper here
  • Guyer, Craig; Mark A. Bailey, and Robert H. Mount 2018. Lizards and snakes of Alabama. University of Alabama Press, 397 pp. - get paper here
  • Harlan, R. 1827. Genera of North American Reptilia, and synopsis of the species. J. Acad. Nat. Sci. Philadelphia 5: 317-372 - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Hart, Sean;Manchester, Sharon 1993. Elaphe guttata emoryi (Great Plains rat snake). USA: Colorado. Herpetological Review 24 (4): 156 - get paper here
  • Hecht-Kardasz, K. A. 2012. Geographic distribution: Pantherophis guttatus (red cornsnake). Herpetological Review 43: 622 - get paper here
  • Hedges SB, Powell R, Henderson RW, Hanson S, and Murphy JC 2019. Definition of the Caribbean Islands biogeographic region, with checklist and recommendations for standardized common names of amphibians and reptiles. Caribbean Herpetology 67: 1–53
  • Hertz, A. 2006. Ein Fall von Kannibalismus bei Pantherophis guttatus. Elaphe 14 (1): 43
  • Hilger, Axel & Axel Kwet 2015. Die Schlangenpest. Terraria-Elaphe 2015 (5): 78-81 - get paper here
  • HIll, Robert L. 2011. Geographic Distribution: Pantherophis guttatus (red cornsnake). Herpetological Review 42 (3): 395 - get paper here
  • Hoehfurtner, T., Wilkinson, A., Nagabaskaran, G., & Burman, O. H. 2021. Does the provision of environmental enrichment affect the behaviour and welfare of captive snakes? Applied Animal Behaviour Science, 239, 105324 - get paper here
  • Hoehfurtner, T., Wilkinson, A., Walker, M., & Burman, O. 2021. Does Enclosure Size Influence the Behaviour & Welfare of Captive Snakes (Pantherophis guttatus)? Applied Animal Behaviour Science, 105435 - get paper here
  • Irwin, Kelly J. 2004. Arkansas Snake Guide. Arkansas Game & Fish Commission, Little Rock, 50 pp.
  • Jasker, M. 2005. Care sheet Pantherophis guttatus emoryi (male). Litteratura Serpentium 25 (1): 46-47 - get paper here
  • Jasker, M. 2006. Caresheet Pantherophis guttatus emoryi (Female). Litteratura Serpentium 26 (3): 214-215 - get paper here
  • Jensen, John B.; Carlos D. Camp, Whit Gibbons, & Matt J. Elliott 2008. Amphibians and reptiles of Georgia. University of Georgia Press, 575 pp.
  • Katti C, Stacey-Solis M, Coronel-Rojas NA and Davies WIL 2019. The Diversity and Adaptive Evolution of Visual Photopigments in Reptiles. Front. Ecol. Evol. 7:352 - get paper here
  • Köhler, G. & Berg, P. 2005. Kornnattern. Herpeton Verlag, Offenbach, 142 pp.
  • Kravchuk, Lindsay; Charles M. Watson 2020. Corn Snakes Can Behaviorally Enhance Crypsis by Choosing Complex Backgrounds and Substrate. Animal Behavior and Cognition 7(1): 39-48 - get paper here
  • Kreutz, R. 2005. Farb- und Zeichnungsstandard der Kornnatter (Panterhophis guttatus). Kirschner und Seufer Verlag, Keltern-Weiler, 158 pp.
  • Kunz, K. 2004. Die Kornnatter (Pantherophis guttatus). Natur und Tier Verlag (Münster), 64 pp. - get paper here
  • Kunz, K. 2007. FAQ Kornnattern. Reptilia (Münster) 12 (65): 87-90 - get paper here
  • Kunz, K. 2015. Dauerbrenner im Terrarium: Nattern. Reptilia (Münster) 20 (115): 18-27 - get paper here
  • Laita, Mark 2013. Serpentine. Abrams and PQ Blackwell, Auckland, New Zealand, 200 unnumbered pages
  • Lannutti, Dominic I.;LaDuc, Travis J.;Johnson, Jerry D. 1996. Geographic Distribution. Elaphe guttata emoryi. Herpetological Review 27 (4): 212 - get paper here
  • Linné, C. von [= Linnaeus, C.] 1766. Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio duodecima, reformata. Laurentii Salvii, Stockholm, Holmiae. 1-532 pp. - get paper here
  • Linzey, D.W., & Clifford, M.J. 1981. Snakes of Virgina. Univ. Press of Virginia, Charlottesville 159 pp.
  • Lönnberg, Einar 1894. Notes on reptiles and batrachians collected in Florida in 1892 and 1893. Proc. US Natl. Mus. 17 (1003): 317-339 - get paper here
  • Love, B. 2011. Okeetee. Reptilia (Münster) 16 (89): 14-15 - get paper here
  • Love, K. 2011. Okeetee Rising - Mein Vierteljahrhundert in der Welt der Kornnattern. Reptilia (Münster) 16 (92): 35-39 - get paper here
  • Love, K. & B. Love 2011. Den Guppys auf der Spur - 50 Jahre Zucht der Kornnatter, Pantherophis guttatus. Reptilia (Münster) 16 (92): 20-27 - get paper here
  • Love, K. & B. Love 2011. Kornnattern vermehren - Ein Rezept für Jedermann. Reptilia (Münster) 16 (92): 28-34 - get paper here
  • Lynch, J.D. 1985. Annotated checklist of the amphibians and reptiles of Nebraska. Nebraska Acad. of Sci., Lincoln, Trans. 13: 33-57.
  • Marques R, Mebert K, Fonseca É, Rödder D, Solé M, Tinôco MS 2016. Composition and natural history notes of the coastal snake assemblage from Northern Bahia, Brazil. ZooKeys 611: 93-142. doi: 10.3897/zookeys.611.9529 - get paper here
  • Marques, R., D. Rödder, M. Solé & M. S. Tinôco 2017. Diversity and habitat use of snakes from the coastal Atlantic rainforest in northeastern Bahia, Brazil. Salamandra 53 (1): 34-43 - get paper here
  • McFadden, Michael S.; Philip Topham, and Peter S. Harlow 2017. A Ticking Time Bomb: Is the illegal pet trade a pathway for the establishment of Corn Snake (Elaphe guttata) populations in Australia? Australian Zoologist In-Press. - get paper here
  • Melo-Dias M., Souza-Cruz P.G., Moreira I.G., Curi N.H.A., Carvalho N.S., Freitas M.A., Rosa C. 2023. Invasive amphibians and reptiles living in Brazil. South American Journal of Herpetology 29: 38–65 - get paper here
  • Merker, C. & Hanley, S. 2005. Pantherophis guttatus - Corn snake. Reptilia (GB) (38): 43-46 - get paper here
  • Miller, Brian T. and Joyce L. Miller. 2010. Geographic distribution: Pantherophis guttatus. Herpetological Review 41 (3): 381-382 - get paper here
  • MIller, Karl E. and Anna Fasoli. 2015. Pantherophis guttatus (red cornsnake) diet. Herpetological Review 46 (1): 105-106 - get paper here
  • Mitchell, J. C. & Reay, K.K. 1999. Atlas of amphibians and reptiles in Virginia. Specialty Publication 1, VA Dept of Game and Fisheries, 122 pp. - get paper here
  • Mitchell, J.C. 1979. The concept of phenology and its application to the study of amphibian and reptile life histories. Herpetological Review 10 (2): 51-54 - get paper here
  • Mitchell, J.C. 1994. The reptiles of Virginia. Virginia Department of Game and Inland Fisheries, ca. 350 pp.
  • MO, MATTHEW & ELOUISE MO. 2021. An amelanistic Red Cornsnake (Pantherophis guttatus) as a possible identity for an unusual road-killed snake discovered in Sydney, Australia. Reptiles & Amphibians 28(3): 480–482. - get paper here
  • Montgomery, Chad;Compton, Leeann;Childers, Theresa;Bergman, Enoch;Manzer, Jerry D.;Sifert, James;Hill, Ben;Mackessy, Stephen P. 1998. Geographic Distribution. Elaphe guttata emoryi. Herpetological Review 29 (2): 112 - get paper here
  • Ollonen, J., Khannoon, E. R., Macrì, S., Vergilov, V., Kuurne, J., Saarikivi, J., ... & Di-Poï, N. 2024. Dynamic evolutionary interplay between ontogenetic skull patterning and whole-head integration. Nature Ecology & Evolution, 1-16 - get paper here
  • O’Shea, M. 2018. The Book of Snakes. Ivy Press / Quarto Publishing, London, - get paper here
  • Paden, L; Ondich, B L; Norton, T M & Steen, D A; 2019. Pantherophis guttatus (Red Cornsnake) Attempted predation and defensive behavior. Herpetological Review 50 (2): 397 - get paper here
  • Palmer, W.M. & Braswell, A.L. 1995. Reptiles of North Carolina. Univ. North Carolina Press
  • Penning, D. A. & Cairns, S. 2012. Growth rates of neonate red cornsnakes, Pantherophis guttatus (Colubridae), when fed in mutally exclusive mass-ratio feeding categories. Herpetological Review 43: 605-607 - get paper here
  • Penning, David A. and Stefan Cairns 2016. Prey-Handling Behaviors of Naïve Pantherophis guttatus. Journal of Herpetology 50 (2): 196-202 - get paper here
  • Petzing, John; Phillips, Christopher A. 1998. Geographic Distribution. Elaphe guttata. Herpetological Review 29 (2): 112 - get paper here
  • Powell, R. & Henderson, R.W. 2003. A second set of addenda to the checklist of West Indian Amphibians ad Reptiles. Herpetological Review 34 (4): 341-345 - get paper here
  • Pyron, R.Alexander; Frank T. Burbrink 2009. Neogene diversification and taxonomic stability in the snake tribe Lampropeltini (Serpentes: Colubridae). Molecular Phylogenetics and Evolution 52 (2): 524-529 - get paper here
  • Questel K, Vitri A. 2012. Pantherophis guttatus (Corn Snake). Distribution. Caribbean Herpetology 37: 1 - get paper here
  • Quijano, Fernando Mendoza; Avila, Ricardo Reyes; Narvaez, Laura Contreras 1998. Geographic Distribution. Elaphe guttata meahllmorum. Herpetological Review 29 (2): 112 - get paper here
  • RODRIGUES NEHEMY, IBRAHIM KAMEL; THAYLLON ORZECHOWSKY GOMES, FERNANDA PAIVA, WESLEY KAUAN KUBO, JOÃO EMÍLIO DE ALMEIDA JÚNIOR, NATHAN FERNANDES NEVES & VINICIUS DE AVELAR SÃO PEDRO. 2022. Herpeto-commerce: A look at the illegal online trade of amphibians and reptiles in Brazil. Cuadernos de Herpetología 36(2): 185–196. - get paper here
  • Rouwkema, J. 2005. Vervelling van Pantherophis guttatus (de Rode rattenslang) - een elektronenmicroscopische studie. Lacerta 63 (5): 191-198 - get paper here
  • Rush, S.A. 2018. Geographic Distribution: Pantherophis guttatus (Red Cornsnake). Herpetological Review 49 (1): 78. - get paper here
  • Rush, Scott A.; Kim Sash, John Carroll, Bill Palmer, and Aaron T. Fisk 2014. Feeding Ecology of the Snake Community of the Red Hills Region Relative to Management for Northern Bobwhite: Assessing the Diet of Snakes Using Stable Isotopes. Copeia Jun 2014, Vol. 2014, No. 2: 288-296. - get paper here
  • Saenko S.V., Lamichhaney S., Martinez Barrio A., Rafati N., Andersson L. & M. C. Milinkovitch 2015. Amelanism in the corn snake is associated with the insertion of an LTR-retrotransposon in the OCA2 gene . Scientific Reports 5, 17118 - get paper here
  • Schmid, A. 2012. Die Entstehung einer neuen Designermorphe der Kornnatter (Pantherophis guttatus). Reptilia (Münster) 17 (95): 42-46 - get paper here
  • Schmidt, D. 2000. Kornnattern und Erdnattern. Natur und Tier Verlag (Münster), 200 pp. - get paper here
  • Schmidt, D. 2004. Zur Reproduktion von Schlangen im Terrarium: Aufgaben - Fakten - Probleme. Draco 5 (17): 22-36 - get paper here
  • Schmidt, D. 2015. Nattern. Arten und Tipps für Einsteiger. Reptilia (Münster) 20 (115): 28-41 - get paper here
  • Schmidt, W. 2002. Kornnattern. Natur und Tier Verlag (Münster), 88 pp. - get paper here
  • Schnitzler, R. 1990. Elaphe guttata (LINNAEUS, 1766). Sauria 12 Suppl.: 161–168 - get paper here
  • Schulz, K. D. 1996. Eine Monographie der Schlangengattung Elaphe Fitzinger. Bushmaster, Berg (CH): 1-460
  • Schulz, Klaus-Dieter 1996. A monograph of the colubrid snakes of the genus Elaphe Fitzinger. Koeltz Scientific Books, 439 pp.
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Shaw, G. 1802. General Zoology, or Systematic Natural History. Vol.3, part 2. G. Kearsley, Thomas Davison, London: 313-615 - get paper here
  • Skubowius, B. 2010. New Jersey – Mit „Fieldherpern“ und „Pineys“ auf der Suche nach der Nördlichen Kiefernnatter (Pituophis melanoleucus melanoleucus). Reptilia (Münster) 15 (84): 52-60 - get paper here
  • SKUBOWIUS, Bernd 2012. Die Lokalvarianten der Kornnatter (Pantherophis guttatus). Terraria-Elaphe 2012 (6):16-25 - get paper here
  • Smith, H.M., D. Chiszar, J.R. Staley & K. Tepedelen 1994. Population relationships in the corn snake Elaphe guttata (Reptilia: Serpentes). Texas Journal of Science 46 (3): 259-292. - get paper here
  • Soehren, Eric C. 2014. Geographical Distribution: Pantherophis guttatus (red cornsnake). Herpetological Review 45 (4): 665 - get paper here
  • Stebbins,R.C. 1985. A Field Guide to Western Reptiles and Amphibians, 2nd ed. Houghton Mifflin, Boston
  • STEVENSON, DIRK J. et al. 2021. New County Records for Amphibians and Reptiles in Georgia, USA. Herpetological Review 52 (2): 350-359
  • Taylor,E.H. 1949. A preliminary account of the herpetology of the state of San Luis Potosi, Mexico. Univ. Kansas Sci. Bull. 33 (2): 169-215 - get paper here
  • Tennant, A. 2003. Snakes of North America - Eastern and Central Regions, revised edition. Lone Star Books, 605 pp.
  • Tennant, A. & Bartlett, R.D. 2000. Snakes of North America - Eastern and Central Regions. Gulf Publishing, Houston, TX, 588 pp.
  • Thawley, Christopher J. and Fern Graves. 2014. Geographical Distribution: Pantherophis guttatus (red cornsnake). Herpetological Review 45 (4): 665 - get paper here
  • Thawley, Christopher J., Mark W. Herr and Sean P. Graham. 2016. Pantherophis guttatus (Red Cornsnake) Defensive behavior. Herpetological Review 47 (1): 151-152 - get paper here
  • Thorpe, R.S. 2022. Reptiles of the Lesser Antilles. Chimaira, Frankfurt, 608 pp. - get paper here
  • Trutnau, L. 1985. Erfahrungen mit der Kornnatter Elaphe guttata (LINNAEUS 1766). Herpetofauna 7 (38): 6-10 - get paper here
  • Ullate-Agote A., Milinkovitch M.C. & A.C. Tzika 2015. The genome sequence of the corn snake (Pantherophis guttatus), a valuable resource for EvoDevo studies in squamates. Int. J. Dev. Biol. 58: 881-888 - get paper here
  • Utiger, Urs, Notker Helfenberger, Beat Schätti, Catherine Schmidt, Markus Ruf and Vincent Ziswiler 2002. Molecular systematics and phylogeny of Old World and New World ratsnakes, Elaphe Auct., and related genera (Reptilia, Squamata, Colubridae). Russ. J. Herpetol. 9 (2): 105-124. - get paper here
  • Van Dorp, D. 2003. Het kweken met Elaphe guttata. Lacerta 61 (2): 73-77 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Werler, John E. & James R. Dixon 2000. Texas Snakes. University of Texas Press, 544 pages
  • Werning, H. 2009. Florida - ein kleiner herpetologischer Reiseführer. Draco 10 (37): 4-23 - get paper here
  • Woodbury, Angus M.;Woodbury, Dixson M. 1942. Studies of the rat snake, Elaphe laeta, with description of a new subspecies. Proc. Biol. Soc. Washington 55: 133-142 - get paper here
 
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