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Pareas formosensis (VAN DENBURGH, 1909)

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Higher TaxaPareatidae, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesFormosa Slug Snake 
SynonymAmblycephalus formosensis VAN DENBURGH 1909
Psammodynastes compressus OSHIMA 1910
Pareas formosensis — HU et al. 1980
Pareas formosensis — GUO & DENG 2008
Pareas formosensis — GUO et al. 2011 
DistributionTaiwan

Type locality: Kanshirei, Formosa Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
TypesHolotype: CAS 18006
Holotype: NSMT H02567 [compressus] 
CommentSynonymy partly after ZHAO & ADLER 1993, Jiang (2004) and Guo & Deng 2009. Amblycephalus komaii MAKI 1931 has been removed from the synonymy of P. formosensis and revalidated by YOU et al. 2015.

Similar species: Pareas formosensis and P. chinensis share very similar coloration and scale patterns (Jiang 2004). However, our molecular phylogenetic results indicate that the two species are not especially closely related within Pareas. Pareas formosensis is the sister species to P. hamptoni, and P. chinensis the sister of P. boulengeri. Close examination of maxillary teeth is consistent with this result. Both P. formosensis and P. hamptoni have 7–8 maxillary teeth, while P. chinensis and P. boulengeri have only 4–5 maxillary teeth (Rao and Yang, 1992). 
EtymologyNamed after its distribution in Taiwan (= Formosa). 
References
  • Barbour, Thomas 1912. Some Chinese Vertebrates: Amphibia and Reptilia. Memoirs of the Museum of Comparative Zoölogy 40 (4): 125-136 - get paper here
  • Guo, Keji & Deng, Xuejiang 2009. A new species of Pareas (Serpentes: Colubridae: Pareatinae) from the Gaoligong Mountains, southwestern China. Zootaxa 2008: 53-60 - get paper here
  • GUO, YUHONG;, YUNKE WU,, SHUNPING HE, HAITAO SHI & ERMI ZHAO 2011. Systematics and molecular phylogenetics of Asian snail-eating snakes (Pareatidae). Zootaxa 3001: 57–64 - get paper here
  • Hu, BU-QING, MET-HUA HUANG, ZHAN-TAI XIE, ER-MI ZHAO, YAO-MING JIANG, QING-YUN HUANG, Yu ZONG, and J 1980. Atlas of Chinese Snakes [in Chinese]. Shanghai Press Sci. Technol., Shanghai. (3), 5, 3, 166 pages
  • Inger R F; Zhao E; Bradley Shaffer H; Guanfu Wu 1990. Report on a collection of amphibians and reptiles from Sichuan, China. Fieldiana: Zoology (No. 58) : i-iii, 1-24 - get paper here
  • JIANG, Yaoming 2004. Pareas chinensis (Babour, 1912) should be a junior synonym of Pareas formosensis (Van Denburgh, 1909). Sichuan Journal of Zoology, 23(3), 209–210 - get paper here
  • Maki, M. 1931. Monograph of the Snakes of Japan. Dai-ichi Shobo, Tokyo. (1), 7, 240 pages (expanded edition 1933)
  • Mell, Rudolf 1922. Beiträge zur Fauna Sinica. i. Die Vertebraten Südchinas; Feldlisten und Feldnoten der Säuger, Vögel, Reptilien, Batrachier. Archiv für Naturgeschichte 88 (10): 1-134
  • Ota H. Lin J T. Hirata T. Chen S L. 1997. Systematic review of Colubrid snakes of the genus Pareas in the East Asian islands. Journal of Herpetology 31 (1): 79-87. - get paper here
  • Rao, D.Q. & D.T. YANG 1992. Phylogenetic systematics of Pareinae (Serpents) of southeastern Asia and adjacent islands with relationship between it and the geology changes. [In Chinese, English summary]. Acta zool. Sinica, 38 (2): 139-150.
  • Stejneger, L. 1910. The batrachians and reptiles of Formosa. Proc. US Natl. Mus. 38: 91-114 - get paper here
  • Toriba, M. 1993. Present location of the type specimens described by Maki. The Snake 25: 71-72
  • Van DENBURGH, JOHN 1909. New and previously unrecorded species of reptiles and amphibians from the island of Formosa. Proc. Cal. Acad. Sci. (4) 3 (3): 49-56. - get paper here
  • You, Chung-Wei; Nikolay A. Poyarkov Jr and Si-Min Lin 2015. Diversity of the snail-eating snakes Pareas (Serpentes, Pareatidae) from Taiwan. Zoologica Scripta, DOI: 10.1111/zsc.12111 - get paper here
  • Zhao,E. & Adler,K. 1993. Herpetology of China. SSAR, Oxford/Ohio, 1-522
 
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