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Philodryas erlandi LÖNNBERG, 1902

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Miranda Green Racer 
SynonymPhilodryas Erlandi LÖNNBERG 1902
Philodryas ternetzii erlandi — MÜLLER 1928
Philodryas mattogrossensis — THOMAS 1976 (part.)
Philodryas mattogrossensis — CEI 1993 (part.)
Philodryas mattogrossensis — LOBO & SCROCCHI 1994
Philodryas mattogrossensis — MORO 1999
Philodryas mattogrossensis — ZAHER 1999
Philodryas mattogrossensis — ARZAMENDIA & GIRAUDO 2002
Philodryas mattogrossensis — KACOLIRIS et al. 2006
Philodryas mattogrossensis — SCHALK 2010
Philodryas erlandi — CACCIALI et al. 2016
Philodryas erlandi — KWET 2017
Philodryas erlandi — MELO-SAMPAIO et al. 2020 
DistributionParaguay, SE Bolivia, N Argentina (Salta, Formosa, Chaco, Santiago del Estero)

Type locality: Tarija, Bolivia  
Reproductionoviparous 
TypesSyntypes: NRM 5097, BMNH 1946.1.8.43 
DiagnosisDiagnosis: Philodryas erlandi differs from all congeners except P. mattogrossensis by a dorsal colour that shifts from green on the anterior half of the body to brown on the posterior part. Philodryas erlandi is distinguished from P. mattogrossensis by the absence of a well-marked dark green or brown dorsal stripe restricted to the anterior one- third of the body (vs present in P. mattogrossensis), pres- ence of a dark line on the supralabials (vs absent or less evident in P. mattogrossensis), dorsal formula usually 19–19–15 (vs usually 21–19–15 in P. mattogrossensis), and first temporal elongated (vs. as wide as high in P. mattogrossensis).
 
CommentSynonymy after CACCIALI et al. 2016.

Distribution: see map in Cacciali et al. 2016: Fig. 11. 
EtymologyNamed in honor of Nils Erland Herbert Nordenskiöld (1877-1932), Swedish archeologist and anthropologist. 
References
  • Arzamendia, V. & A. R. Giraudo 2002. Lista y distribución de los ofidios (Reptilia: Serpentes) de Santa Fe, Argentina. Cuadernos de Herpetología 16: 15–32
  • Cacciali, P., H. Cabral, V. L. Ferreira & G. Köhler 2016. Revision of Philodryas mattogrossensis with the revalidation of P. erlandi (Reptilia: Squamata: Dipsadidae). Salamandra 52 (4): 293-305 - get paper here
  • Cei, J. M. 1993. Reptiles del noroeste, nordeste y este de la Argentina. Museo Regionale Sci. Naturale Torino, Monografie 14: 1-949
  • Kacoliris F.P.; Berkunsky I. & Williams J. 2006. Herpetofauna of Impenetrable, Argentinean Great Chaco. Phyllomedusa 5 (2): 149-158 - get paper here
  • Kwet, Axel 2017. Neue Arten: Liste der im Jahr 2016 neu beschriebenen Reptilien. Terraria-Elaphe 2017 (3): 54-70 - get paper here
  • Lobo, F. & G. Scrocchi 1994. Osteología craneal del género Philodryas (Serpentes: Colubridae). Cuadernos de Herpetología 8: 104–111
  • Lönnberg, E. 1902. On a collection of snakes from north-western Argentina and Bolivia containing new species. Ann. Mag. nat. Hist. (7) 10: 457-462 - get paper here
  • Melo-Sampaio, P. R., Passos, P., Martins, A. R., Moura-leite, J. C., Morato, S. A., Venegas, P. J., Chávez G., Venâncio N.M. & De Souza, M. B. 2020. A phantom on the trees: integrative taxonomy supports a reappraisal of rear-fanged snakes classification (Dipsadidae: Philodryadini). Zoologischer Anzeiger 290: 19-39 [dated 2021 in print version, but 2020 published online] - get paper here
  • Moro, S. 1999. Relaciones filogenéticas de Xenodontinae (Ser- pentes) neotropicales basadas en musculatura craneal. Cuadernos de Herpetología 13: 19–35
  • Müller, L. 1928. Herpetologische Mitteilungen. Zool. Anz. 77: 61-84
  • Schalk, Christopher M. 2010. Philodryas mattogrossensis: Diet. Herpetological Review 41 (3): 372 - get paper here
  • Zaher, H. 1999. Hemipenial morphology of the South American xenodontine snakes, with a proposal for a monophyletic Xenodontinae and a reappraisal of colobrid hemipenes. Bull. Amer. Mus. Nat. Hist., 240: 1-168 - get paper here
 
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