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Philodryas mattogrossensis KOSLOWSKY, 1898

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Miranda Green Racer
Portuguese: Cobra-Cipó, Cobra-do-Papo-Amarelo 
SynonymPhilodryas mattogrossensis KOSLOWSKY 1898: 29
Philodryas ternetzii SCHENKEL 1901
Philodryas boulengeri WENER 1909
Chlorosoma mattogrossensis — AMARAL 1929
Philodryas mattogrossensis — PETERS & OREJAS-MIRANDA 1970
Philodryas mattogrossensis — CEI 1993
Philodryas mattogrossensis — LEYNAUD & BUCHER 1999: 29
Philodryas mattogrossensis — BERNILS et al. 2007
Philodryas mattogrossensis — GRAZZIOTIN et al. 2012
Philodryas mattogrossensis — WALLACH et al. 2014: 551
Philodryas mattogrossensis — CACCIALI et al. 2016
Philodryas mattogrossensis — NOGUEIRA et al. 2019
Philodryas mattogrossensis — MELO-SAMPAIO et al. 2020 
DistributionSW Brazil (Mato Grosso, Goias), Paraguay, Argentina (Salta, NW Santiago del Estero)

Type locality: Miranda, Mato Grosso, Brazil (the state of Mato Grosso was subdivided and Miranda is now in the Mato Grosso do Sul state). Neotype locality: “Taunay, Mato Grosso do Sul (-20.2858° S, -56.0847° W), Brazil”

ternetzii: Type locality: “Bemalcue” in Paraguay, actually “Bernal Cué” (Departamento Cordillera, Paraguay) (Cacciali & Wuest 2009).  
Reproductionoviparous 
TypesNeotype: SMF 49990 (designated by Cacciali et al. 2016); Holotype: MLP, an 1195 mm female (C. Bach), now lost (Ferraro & Williams 2006).
Holotype: NMBA 1939 [ternetzii] 
Diagnosis 
CommentSynonymy after AMARAL 1929, PETERS & OREJAS-MIRANDA 1970, CEI 1993, and CACCIALI et al. 2016. Philodryas Erlandi LÖNNBERG 1902 was removed from the synonymy of Philodryas mattogrossensis by CACCIALI et al. 2016.

Distribution: see map in Cacciali et al. 2016: Fig. 11. See map in Nogueira et al. 2019.

Habitat: partly arboreal (Harrington et al. 2018). 
EtymologyNamed after the type locality. 
References
  • Amaral,A. 1977. Serpentes do Brasil. Iconografía colorida. Tomo I, II y III. Ed. Melhoramentos & Inst. Nac. Livro, Edit. Univ. Sao Paulo, 248 pp.
  • ARAUJO, C.O. & ALMEIDA-SANTOS, S.M. 2011. Herpetofauna de um remanescente de Cerrado no estado de São Paulo, sudeste do Brasil. Biota Neotrop. 11(3): 47-62 - get paper here
  • Bressan, Paulo Magalhães; Maria Cecília Martins Kierulff, Angélica Midori Sugieda (eds.) 2009. FAUNA AMEAÇADA DE EXTINÇÃO NO ESTADO DE SÃO PAULO: VERTEBRADOS. São Paulo: Fundação Parque Zoológico de São Paulo: Secretaria do Meio Ambiente, 2009. - get paper here
  • Cacciali, P., H. Cabral, V. L. Ferreira & G. Köhler 2016. Revision of Philodryas mattogrossensis with the revalidation of P. erlandi (Reptilia: Squamata: Dipsadidae). Salamandra 52 (4): 293-305 - get paper here
  • Cacciali, Pier; Norman J. Scott, Aida Luz Aquino Ortíz, Lee A. Fitzgerald, and Paul Smith 2016. The Reptiles of Paraguay: Literature, Distribution, and an Annotated Taxonomic Checklist. SPECIAL PUBLICATION OF THE MUSEUM OF SOUTHWESTERN BIOLOGY, NUMBER 11: 1–373 - get paper here
  • Cei, J. M. 1993. Reptiles del noroeste, nordeste y este de la Argentina. Museo Regionale Sci. Naturale Torino, Monografie 14: 1-949
  • Fowler, I.R.; Salomao, M.D. 1995. A new technique to distinguish between immature and adult snakes and between males and females in six species of the neotropical colubrid snakes Philodryas. Studies on Neotropical Fauna and Environment 30 (3): 149-157 - get paper here
  • Giraudo, A. R., and G. J. Scrocchi 2002. Argentinian Snakes: An Annotated Checklist. Smithsonian Herp. Inf. Serv. (132): 1-53 - get paper here
  • Gonzalez R. C. et al. 2020. Lista dos Nomes Populares dos Répteis no Brasil – Primeira Versão. Herpetologia Brasileira 9 (2): 121 – 214 - get paper here
  • Grazziotin, Felipe G.; Hussam Zaher, Robert W. Murphy, Gustavo Scrocchi, Marco A. Benavides, Ya-Ping Zhang and Sandro L. Bonatto 2012. Molecular phylogeny of the New World Dipsadidae (Serpentes: Colubroidea): a reappraisal. Cladistics DOI: 10.1111/j.1096-0031.2012.00393.x - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Kacoliris F.P.; Berkunsky I. & Williams J. 2006. Herpetofauna of Impenetrable, Argentinean Great Chaco. Phyllomedusa 5 (2): 149-158 - get paper here
  • Koslowsky,J. 1898. Ofidios de Matto-Grosso (Brasil). Rev. Mus. La Plata 8: 25-34 - get paper here
  • Kwet, Axel 2017. Neue Arten: Liste der im Jahr 2016 neu beschriebenen Reptilien. Terraria-Elaphe 2017 (3): 54-70 - get paper here
  • Leynaud, Gerardo C. and Enrique H. Bucher 1999. La fauna de serpientes del Chaco Sudamericano: diversidad, distribución geografica y estado de conservación. Academia Nacional de Ciencias Miscelanea (98):1-46
  • MARQUES, O. A. V. et al. 2009. Repteis. In: Bressan et al., FAUNA AMEAÇADA DE EXTINÇÃO NO ESTADO DE SÃO PAULO: VERTEBRADOS. São Paulo: Fundação Parque Zoológico de São Paulo: Secretaria do Meio Ambiente, pp. 285-327 - get paper here
  • Mattison, Chris 2007. The New Encyclopedia of Snakes. Princeton University Press
  • Melo-Sampaio, P. R., Passos, P., Martins, A. R., Moura-leite, J. C., Morato, S. A., Venegas, P. J., Chávez G., Venâncio N.M. & De Souza, M. B. 2020. A phantom on the trees: integrative taxonomy supports a reappraisal of rear-fanged snakes classification (Dipsadidae: Philodryadini). Zoologischer Anzeiger 290: 19-39 [dated 2021 in print version, but 2020 published online] - get paper here
  • Nogueira, Cristiano C.; Antonio J.S. Argôlo, Vanesa Arzamendia, Josué A. Azevedo, Fausto E. Barbo, Renato S. Bérnils, Bruna E. Bolochio, Marcio Borges-Martins, Marcela Brasil-Godinho, Henrique Braz, Marcus A. Buononato, Diego F. Cisneros-Heredia, 2019. Atlas of Brazilian snakes: verified point-locality maps to mitigate the Wallacean shortfall in a megadiverse snake fauna. South American J. Herp. 14 (Special Issue 1):1-274 - get paper here
  • PEIXOTO COUTO, RODNEY MURILLO & MAIARA CABRERA MIGUEL. 2021. PHILODRYAS MATTOGROSSENSIS. DIET. Herpetological Review 52 (3): 674–675. - get paper here
  • Ramalho, Werther Pereira; Vinícius Guerra Batista; Luciano Roberto Passos Lozi 2014. Anfíbios e répteis do médio rio Aporé, estados de Mato Grosso do Sul e Goiás, Brasil. Neotropical Biology and Conservation 9 (3): 147-160; doi: 10.4013/nbc.2014.93.04 - get paper here
  • Scrocchi, G.J.; Moreta, J.C. & Kretzschmar, S. 2006. Serpientes del Noroeste Argento [Jujuy, Salta, Tucumán, Catamarca, La Rioja, Santiago del Estero]. Fundación Miguel Lillo, Tucumán, 178 pp.
  • Vaz-Silva, W. et al. 2007. Herpetofauna, Espora Hydroelectric Power Plant, state of Goiás, Brazil. Check List 3 (4): 338-345 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Weiler, Andrea 2015. Powerful venom in Philodryas mattogrossensis (Serpentes: Dipsadidae) feeding on a Teius teyou (Sauria: Teiidae). Herpetology Notes 8: 545-547 - get paper here
  • Williams,J.D. & Francini,F. 1991. A checklist of the Argentine snakes. Boll. Mus. Reg. Sci. Nat. Torino 9 (1): 55-90
  • Zaher, Hussam; Fausto Erritto BarboI; Paola Sanchez Martínez; Cristiano Nogueira; Miguel Trefaut Rodrigues; Ricardo Jannini Sawaya 2011. Répteis do Estado de São Paulo: conhecimento atual e perspectivas. Biota Neotropica, 11 (1): 1–15. - get paper here
 
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