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Philothamnus thomensis BOCAGE, 1882

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Sao Tome Wood Snake 
SynonymPhilothamnus thomensis BOCAGE 1882: 11
Philothamnus thomensis — BOULENGER 1894: 101
Philothamnus thomensis — GÜNTHER 1895: 528
Philothamnus thomensis — STERNFELD 1917
Philothamnus semivariegatus thomensis — MERTENS 1934
Philothamnus semivariegatus thomensis — LOVERIDGE 1958: 125
Philothamnus thomensis — HUGHES 1985: 524
Philothamnus thomensis — JESUS et al. 2009
Philothamnus thomensis — WALLACH et al. 2014: 559
Philothamnus thomensis — CERÍACO et al. 2023 
DistributionSão Tomé (Gulf of Guinea), Uganda ?

Type locality: "Demarara" (in error)  
Reproductionoviparous 
TypesType: BMNH 1946.1.21.60 (and possibly additional specimens). 
DiagnosisAdditional details, e.g. a detailed description or comparisons (418 characters), are available for collaborators and contributors. Please contact us for details. 
CommentHabitat: in coffee trees, fully arboreal (Harrington et al. 2018)

Distribution: listed for Uganda by Behangana et al. 2023 as P. “thomensis / carinatus”. 
EtymologyNamed after the type locality, São Tomé. 
References
  • Behangana, M., Ayebare, S., Magala, R., Lukwago, W., Ssekisambu, R. S., Alezuyo, C., ... & Hughes, D. 2023. Establishing baseline data for herpetofauna in western Uganda, an East African biodiversity hotspot: Reptile and amphibian diversity in Uganda. Herpetology Notes, 16, 773-789
  • Bocage, J.V.B. de 1882. Notice sur les espèces du genre "Philothamnus" qui se trouvent au Muséum de Lisbonne. Jorn. Sci. Math. Phys. Nat. Lisboa 9 (33): 1-20 - get paper here
  • Boulenger, George A. 1894. Catalogue of the snakes in the British Museum (Natural History). Volume II., Containing the Conclusion of the Colubridæ Aglyphæ. British Mus. (Nat. Hist.), London, xi, 382 pp. - get paper here
  • Ceríaco, L. M. P., Marques, M. P., de Sousa, A. C. A., Veríssimo, J., Beja, P., & Ferreira, S. 2023. Illustrated keys and a DNA barcode reference library of the amphibians and terrestrial reptiles (Amphibia, Reptilia) of São Tomé and Príncipe (Gulf of Guinea, West Africa). ZooKeys 1168: 41-75
  • Chippaux, Jean-Philippe & Kate Jackson 2019. Snakes of Central and Western Africa. Johns Hopkins University Press, 448 pp. [detaileld review in HR 51 (1): 161] - get paper here
  • Engelbrecht, H.M., Branch, W.R., Greenbaum, E., Alexander, G.J., Jackson, K., Burger, M., Conradie, W., Kusamba, C., Zassi-Boulou, A-G., Tolley, K.A. 2018. Diversifying into the branches: species boundaries in African green and bush snakes, Philothamnus (Serpentes: Colubridae). Molecular Phylogenetics and Evolution 130: 357-365 - get paper here
  • Günther, A. 1895. Notice of Reptiles and Batrachians collected in the eastern half of tropical Africa. Ann. Mag. nat. Hist. (6) 15: 523-529. - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Hughes, B. 1985. Progress on a taxonomic revision of African green tree snakes (Philothamnus spp.). In: Schuchmann, K.L. (ed.) Proceedings of the International Symposium on African Vertebrates. ZFMK, Bonn, pp. 511-530
  • Jesus, J.; Nagy, Z.; Branch, W.; Wink, M.; Brehm, A. & Harris, D. J. 2009. Phylogenetic relationships of African green snakes (genera Philothamnus and Hapsidophrys) from São Tomé, Príncipe and Annobon islands based on mtDNA sequences and comments on their colonization and taxonomy. The Herpetological Journal 19: 41-48 - get paper here
  • Loveridge,A. 1958. Revision of five african snake genera. Bull. Mus. Comp. Zool. Harvard 119: 1-198 (141) - get paper here
  • Meirte, D. 1992. Clés de détermination des serpents d'Afrique. Museum Royal d'Afrique Centrale, Tervuren Belgique Annual Series Octavo Science Zoologique 267:1-152
  • Mertens, R. 1934. Die Inseln-Reptilien, ihre Ausbreitung Variation und Artbildung. Zoologica (Stuttgart) 32: 1-209.
  • Pollo, César J. 2017. Guide of Amphibians and Reptiles of São Tomé and Príncipe. Smashwords / César J. Pollo, 156 pp. - get paper here
  • Sternfeld, R. 1917. Reptilia und Amphibia. In: Schubotz, H. (Hrsg.): Wissenschaftliche Ergebnisse der Zweiten Deutschen Zentral-Afrika-Expedition, 1910-1911 unter Führung Adolph Friedrichs, Herzog zu Mecklenburg. Leipzig: Klinkhardt & Biermann, [Band] 1, Zoologie, Lieferung 11; S. 407-510. - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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