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Phoxophrys tuberculata HUBRECHT, 1881

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Hubrecht's Eyebrow Lizard 
SynonymPhoxophrys tuberculata HUBRECHT 1881: 51
Phoxophrys tuberculata — BOULENGER 1885: 280
Phoxophrys tuberculata — DE ROOIJ 1915: 94
Japalura robinsoni BOULENGER 1920: 287 (fide INGER 1960)
Phoxophrys tuberculata — INGER 1960: 225
Phoxophrys tuberculata — WERMUTH 1967: 75
Phoxophrys tuberculata — MANTHEY & GROSSMANN 1997: 197
Phoxophrys tuberculata — TEYNIÉ et al. 2010
Phoxophrys tuberculata — HARVEY et al. 2019 
DistributionIndonesia (W Sumatra: Jambi, Sumatera Barat, and Sumatera Utara Provinces)

Type locality: Batang, Singalang, West Sumatra.  
Reproductionoviparous 
TypesType: BMNH 1946.8.13.92 (and possibly additional specimens).
Holotype: BMNH [robinsoni] 
DiagnosisDiagnosis (genus). Small draconine agamids reaching a SVL of at least 43 mm and distinguished from all other Agamidae by the following combination of characters: (1) tympanum and external auditory meatus absent, extrastapes attaching to skin; (2) head robust, 77.4–92.1% as wide as long, comprising 27.4–28.5% of SVL; (3) rostral single; (4) prominent tubercular scale positioned below corner of jaw; (5) supercilium projecting upward and outward as pointed, triangular flap; (6) dorsal scales heterogenous with larger tubercular scales interspersed among smaller keeled scales, 56–67 scales around midbody; (7) premaxillary processes of mandible abutting premaxilla; (8) frontal and parietal extensively sculptured, not containing pineal foramen; (9) transverse shorting of prefrontal bone allowing nasal–frontal contact; (10) squamosal extending anteriorly below jugal to curve ventrally and brace against vertical posterior margin of jugal; (11) angular process of mandible absent (Harvey et al. 2019: 94). 
CommentType species: Phoxophrys tuberculata HUBRECHT 1881 is the type species of the genus Phoxophrys HUBRECHT 1881. 
EtymologyNamed after Latin tuberculata, having small swellings. 
References
  • Aguilar-López JL, Luría-Manzano R, Pineda E, Canseco-Márquez L 2021. Selva Zoque, Mexico: an important Mesoamerican tropical region for reptile species diversity and conservation. ZooKeys 1054: 127-153 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Boulenger, G.A. 1920. Results of an expedition to Korinchi Peak 12.400 ft, Sumatra. III. Reptiles & batrachians collected in Korinchi, West Sumatra by Messrs. H.C. Robinson & C. Boden Kloss. J. Federated Malay State Museum, Singapore, 8: 285-296 (other volumes in BHL)
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Denzer, W. 1996. Reptilien und Amphibien der Santubong Halbinsel, Sarawak, Borneo. Sauria 18 (4): 35-42 - get paper here
  • Harvey, Michael B., Larson, Thorton R., Jacobs, Justin L., Shaney, Kyle, Streicher, Jeffrey W., Hamidy, Amir, Kurniawan, Nia & Smit, Eric N. 2019. Phoxophrys After 60 Years: Review of Morphology, Phylogeny, Status of Pelturagonia, and a New Species from Southeastern Kalimantan. Herpetological Monographs 33 (1): 71-107 - get paper here
  • Hubrecht, A.A.W. 1881. On a new genus and species of Agamidae from Sumatra. Notes from the Leyden Museum 3: 51-52. - get paper here
  • Inger, R. F. 1960. A review of the agamid lizards of the genus Phoxophrys HUBRECHT. Copeia 1960 (3): 221 -225. - get paper here
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. (ISBN 3-931587-12-6) - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
 
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