You are here » home advanced search search results Phrynocephalus mystaceus

Phrynocephalus mystaceus (PALLAS, 1776)

Can you confirm these amateur observations of Phrynocephalus mystaceus?

Add your own observation of
Phrynocephalus mystaceus »

Find more photos by Google images search: Google images

Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
SubspeciesPhrynocephalus mystaceus mystaceus (PALLAS 1776)
Phrynocephalus mystaceus khorasanus SOLOVYEVA, DUNAYEV, NAZAROV, RADJABIZADEH & POYARKOV 2018 
Common NamesE: Secret Toadhead Agama
G: Bärtiger Krötenkopf
E: Khorasan Secret Toad-headed Agama [khorasanus]
Russian: Ушастая круглоголовка
Chinese: 大耳沙蜥 
SynonymLacerta mystacea PALLAS 1776: 702
Lacerta aurita PALLAS 1776: Tab. V, fig. 1
Ameiua aurita MEYER 1795: 29 (error typographicus)
Lacerta aurita — SHAW & NODDER 1803: plate 594
Phrynocephalus mystacia — KAUP 1827
Megalochilus auritus — MÜLLER 1881
Phrynocephalus mystaceus — BOULENGER 1885: 379
Phrynocephalus mystaceus mystaceus natio dagestanica KRASSOWSKY 1932
Phrynocephalus mystaceus galli KRASSOWSKY 1932
Megalochilus mystaceus — ANANJEVA 1986
Phrynocephalus mystaceus — WERMUTH 1967: 84
Phrynocephalus mystaceus — ENGELMANN et al 1993
Phrynocephalus mystaceus — ŠMÍD et al. 2014
Phrynocephalus (Megalochilus) mystaceus — SOLOVYEVA et al. 2023

Phrynocephalus mystaceus mystaceus (PALLAS 1776)
Lacerta aurita GMELIN 1789: 1073 (nom. subst. pro Lacerta mystacea)
Gekko auritus — LATREILLE 1801: 61.
Lacerta lobata SHAW 1802: 244 (nom. subst. pro Lacerta aurita)
Agama aurita — DAUDIN 1802: 429
Agama mystacea — MERREM 1820: 53
Agama aurita — LIECHTENSTEIN 1823: 142
Megalochilus auritus — EICHWALD 1831: 185
Phrynocephalus auritus — EVERSMANN 1834: 360
Phrynocephalus auritus — DUMÉRIL & BIBRON 1837: 524
Saccostoma auritum — FITZINGER 1843: 87
Phrynocephalus mystaceus — BOETTGER 1880: 260
Phrynocephalus mystaceus mystaceus — KRASSOWSKY 1932: 227
Phrynocephalus mystaceus Galli KRASSOWSKY 1932: 227
Phrynocephalus mystaceus mystaceus — WERMUTH 1967: 84
Phrynocephalus mystaceus aurantiacaudatus SEMENOV & SHENBROT 1990: 79
Phrynocephalus mystaceus mystaceus — MANTHEY & SCHUSTER 1999: 90
Phrynocephalus (Phrynocephalus) mystaceus mystaceus — BARABANOV & ANANJEVA 2007
Phrynocephalus (Phrynocephalus) mystaceus galli — BARABANOV & ANANJEVA 2007
Phrynocephalus mystaceus aurantiacaudatus — SINDACO & JEREMČENKO 2008
Phrynocephalus mystaceus galli — SINDACO & JEREMČENKO 2008
Phrynocephalus mystaceus galli — MILTO & BARABANOV 2012

Phrynocephalus mystaceus khorasanus SOLOVYEVA, DUNAYEV, NAZAROV, RADJABIZADEH & POYARKOV 2018
Phrynocephalus (Megalochilus) khorasanus — SOLOVYEVA et al. 2023 
DistributionSoviet Union, NW cost of Caspian Sea to E Kazakhstan, Russia (Dagestan, Astrakhan Oblast), Kazakhstan, Turkmenistan, Tajikistan, Uzbekistan, Kyrgyzstan, NE Iran, Afghanistan, NW China (W Xinjiang)

aurantiacaudatus: Kazakhstan
galli: Type locality: Repeteka, Turkestan.

galli: Turkmenistan, Uzbekistan, Kazakhstan, northeastern and eastern Iran and adjacent Afghanistan and Xinjiang, China; Terra typica: “Aus Repeteka andetrifft” [= vicinity of Repetek station, approximately 38º35' N, 63º11' E, Lebapsky Region, Turkmenistan].

khorasanus: Iran (Khorasan Razavi Province); Type locality: Iran, Khorasan historical area, Khorasan Razavi Province (ostan), environs of Gonabad, the right bank of the Kale-Shur River; sand dunes (see Fig. 9 in Solovyeva et al. 2018); N34°39', E58°43'; elevation 850 m.

Type locality: East Caucasus. Restricted to “Naryn-Steppe, an der Nordküste des kaspischen Maeeres” by MERTENS & MÜLLER 1928; Restricted by neotype designation to Ryn Peski (Ryn Sands), Uralskaya Region, north-western Kazakh-
stan, by BARABANOV & ANANJEVA 2007.  
ReproductionOviparous 
TypesNeotype: ZISP 8735.1, adult male, designated by BARABANOV & ANAJEVA 2007
Holotype. ZMMU R-11913 (adult female); paratypes: ZMMU [khorasanus]
Lectotype: ZMMU Re-6413 (designated by Semenov & Shenbrot, 1990: 78) [galli] 
DiagnosisDiagnosis: “Vor den anderen Arten der Gattung durch deutliche querverlaufende Hautfalten im Hals-Nackenbereich ausgezeichnet. Unterscheidet sich von P. mystaceus und P. guttatus durch eine heterogene Dorsalbeschuppung, wobei vergrößerte Schuppen, teils in Gruppen, auf dem Rücken, den Flanken sowie oberseits und seitlich der Schwanzwurzel eingestreut sind. Diese vergrößerten Schuppen heben sich als typische Tuberkeln von der sonst glatten Beschuppung ab. Beiderseits des Nackens liegen paarige große Augenflecken, die bei blauer Umrandung ein rosafarbe­ nes oder rotes Zentrum haben.
Schädel breit, mit steil nach vorn abfallender Schnauzenpartie. Gegenüber dem Schädel von P. mystaceus sind außer den kleineren Abmessungen die höhere Zahnzahl und das relativ größere Foramen parietale charakteri­stisch.” (Ananjeva 1981: 191 who also has a more detailed description).


Additional details (3933 characters) are available for collaborators and contributors. Please contact us for details. 
CommentSynonymy and subspecies mainly after KHALIKOV & ANANJEVA (pers. comm.), WERMUTH 1967, and BARABANOV & ANANJEVA 2007. SEMENOV & SHENBROT 1990 considered galli as a synonym of P. mystaceus. SOLOVYEVA et al. consider P. m. galli as a synonym of mystaceus.

Distribution: See map in SMID et al. 2014 for distribution in Iran.

NCBI taxon ID: 662673 [aurantiacaudatus]
NCBI taxon ID: 662675 [galli]
NCBI taxon ID: 2172839 [khorasanus]
NCBI taxon ID: 662674 [mystaceus]

Etymology (khorasanus). The name of the new subspecies khorasanus is a Latinized toponymic adjective, derived from Khorasan, the name of the historic area and a Khorasan Razavi Province in the northeast Iran, where the new subspecies was found. 
EtymologyNamed after the appearance of this species which is fascinating like a sorcerer; the epithet mystaceus is an masculine adjective formed from suffix L. -ace ‘of the nature of’, hence ‘resembling’ and L. mysta (relating to mystery) meaning ‛a priest of the mysteries and secret rites of divine worship’; mystacea is feminine (Partridge, 1966; Lewis, 1969, cited in Mikaili & Shayegh 2011). However, it could also have been named after Greek “mystax” = beard, moustache :) 
References
  • Ananjeva, N. & Orlov, N. 2005. Lizards of North Eurasia. Reptilia (GB) (38): 54-63 - get paper here
  • Ananjeva, N. B. 1986. O rodovoj samostoyatelnosti ushastoj kriglogolovki, Megalochilus mystaceus (PALLAS 1776). [On the validity of Megalochilus mystaceus (PALLAS 1776).]. Trud. Zool. Inst. AN SSSR, Leningrad 157: 4-13
  • Ananjeva, Natalia B. 2012. Research on Asian Agamid Lizards: Contribution of Zoological Institute, Russian Academy of Sciences, St. Petersburg. Russ. J. Herpetol. 19 (2): 85-100 - get paper here
  • ANANJEVA, Natalia B.; Xianguang GUO and Yuezhao WANG 2011. Taxonomic Diversity of Agamid Lizards (Reptilia, Sauria, Acrodonta, Agamidae) from China: A Comparative Analysis. Asian Herpetological Research 2 (3): 117-128 - get paper here
  • Ananjeva, Natalia B.;Tuniyev, Boris S. 1992. Historical biogeography of the Phrynocephalus species of the USSR. Asiatic Herpetological Research 4: 76-98 - get paper here
  • Ananjeva, Natalija B. 1981. Phrynocephalus mystaceus - Bärtiger Krötenkopf. In: Böhme,W. (ed.), Handbuch der Reptilien und Amphibien Europas, Band I., Echsen I. Akademische Verlagsgesellschaft, Wiesbaden, pp. 203-216
  • Ananjeva, NB, Orlov NL; Khalikov RG; Darevsky IS; Ryabov IS & Barabanov AV 2006. The Reptiles of North Eurasia. Taxonomic Diversity, Distribution, Conservation Status [this comprises the territory of the former Soviet Union and Mongolia]. Pensoft Series Faunistica 47, 250 pp.
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • Anooshe, K., Sohrab, A., Masoud, Y., Eskandar, R. P., Mahdi, R., Faraham, A., ... & Loïc, P. 2020. Reptile species richness associated to ecological and historical variables in Iran. Scientific Reports (Nature Publisher Group), 10(1) - get paper here
  • Bakiev, A., Kirillov, A., Kirillova, N., Ruchin, A., Klenina, A., Gorelov, R., & Kostina, N. 2020. Reptile occurrences data in the Volga River basin (Russia). Biodiversity Data Journal, 8: e58033 - get paper here
  • BARABANOV, A.V. & N.B. ANANJEVA 2007. Catalogue of the available scientific species-group names for lizards of the genus Phrynocephalus Kaup, 1825 (Reptilia, Sauria, Agamidae). Zootaxa 1399: 1-56 - get paper here
  • Bondarenko D. A., Ergashev U. H. 2022. Reptiles of the Southwestern Tajikistan desert plains: Spatial distribution, population density and communities structure. [in Russian] Current Studies in Herpetology 22 (1-2): 17 - get paper here
  • Borkin, L.J. 2022. EARLY STUDIES ON THE HERPETOFAUNA OF KAZAKHSTAN (XVIII CENTURY) [in russian] Almaty: Institute of Zoology of the Republic of Kazakhstan, 2022. - 156 pp.
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Brushko, Z.K. 1995. Lizards of desert regions of Kazakhstan. [in Russian] Kazakhstan National Academy of Sciences, Institute of Zoology, Almaty, 232 pp.
  • Cherlin V. A., Okstein I. L., Aligadzhiev E. 2023. Thermobiological characters of toad-headed agamas (Phrynocephalus mystaceus) (Reptilia, Agamidae) on the Sarykum dune (Dagestan, Russia) obtained by using of temperature loggers. Current Studies in Herpetology 23 (3): 160 - get paper here
  • CHIRIKOVA, M. A., Y. A. ZIMA, M. V. PESTOV, AND V. A. TERENTYEV 2020. Biodiversity of the Herpetofauna of the Muyunkum Desert, Kazakhstan. Herpetological Review 51: 438-446.
  • Davletbakov, A. T.; L. A. Kustareva, D. A. Milko, A. N. Ostastshenko, S. S. Sagymbaev, S. A. Toropov, N. V. Trotchenko 2015. Genetical Fund Cadastre of Kyrgyz Republic. Vol. IV. Phylum Chordata (chordates). Bishkek, 128 p. + XXXVI color tables - get paper here
  • Dieckmann, M. 2011. Bemerkungen zur Haltung des Bärtigen Krötenkopf Phrynocephalus mystaceus (PALLAS 1776). Iguana Rundschreiben 24 (1): 21-26
  • Dujsebayeva, T. N. (ed.) 2010. Short review of last changes in the checklist of amphibians and reptiles of Kazakhstan. [in Russian]. In: Dujsebayeva, T. N. (ed.) Herpetological Researches in Kazakhstan and adjacent countries, p. 37-52
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Dunaev E.A., Orlova V.F. 2017. Amphibians and reptiles of Russia. Atlas and determination. 2nd. ed. Moscow: Phyton XXI, 328 p
  • Dunayev, E. A. 1996. On the possible use of the Ethological features in the taxonomy and phylogeny of toad agamas, Phrynocephalus (Reptilia, Agamidae). Russ. J. Herpetol. 3 (1): 32-38 - get paper here
  • Engelmann, W.E. et al. 1993. Lurche und Kriechtiere Europas. Neumann Verlag (Radebeul, Germany), 440 pp.
  • Eversmann,E. 1834. Lacertae Imperii Rossici variis in itineribus meis observatae. [Lacerta sylvicola]. Nouv. Mém. Soc. imp. Natur. (Moscow) 3: 339-358 - get paper here
  • Gmelin, J.F. 1789. Caroli a Linné Systema naturae. 13. ed., Tom 1 Pars 3. G. E. Beer, Lipsiae. 1033-1516 pp. - get paper here
  • Hoogmoed, M.S. 1980. Herpetologische waarnemingen in Ghana [part 3]. Lacerta 38 (5): 42-48 - get paper here
  • Jablonski D, Regan JM, Holzheuser C, Farooqi J, Basit A, Masroor R 2019. Additional data to the herpetofauna of Afghanistan. Herpetozoa 32: 177-193 - get paper here
  • Kabisch, K. & Belter, H. 1968. Das Verzehren von Amphibien und Reptilien durch Vögel. Zoologische Abhandlungen aus dem Staatlichen Museum für Tierkunde in Dresden 29: 191-227
  • Kamali, Kamran 2020. A Field Guide to the Reptiles and Amphibians of Iran. Edition Chimaira, Frankfurt am Main, Germany (www.chimaira.de). 574 pp.
  • Krassowsky, D.B. 1932. Beitrag zur Systematik von Phrynocephalus mystaceus. Zool. Anz. 97: 225-228
  • Kukushkin O.V., Doronin I.V., Tuniyev B.S., Ananjeva N.B., Doronina M.A. 2017. Introduction of Amphibians and Reptiles at the Caucasus and the Crimea: an Overview and Some Actual Data. (in Russian). Current Studies in Herpetology 17 (3-4): 157–197 . DOI: 10.18500/1814-6090-2017-17-3-4-157-197 - get paper here
  • Kwet, A. 2019. Liste der im Jahr 2018 neu beschriebenen Reptilien. Elaphe 2019 (3): 52-72
  • Lambert, M.R.K. 2002. Preliminary observations on herpetofaunal diversity in the Almaty region, Southern Kazakhstan (September 1998). Herpetological Bulletin (79): 7-13 - get paper here
  • Leviton, Alan E. 1959. Report on a collection of reptiles from Afghanistan. Proc. Cal. Acad. Sci. 29 (12): 445-463 - get paper here
  • Lichtenstein 1823. Amphibien. in: Evermann, Reise nach Buchara: 144. - get paper here
  • Macey, J. R., J. A. Schulte II, A. Larson, N. B. Ananjeva, Y. Wang, R. Pethiyagoda, N. Rastegar-Pouyani, T. J. Papenfuss 2000. Evaluating trans-Tethys migration: an example using acrodont lizard phylogenetics. Systematic Biology 49 (2): 233-256 - get paper here
  • Manthey,U. & SCHUSTER,N. 1999. Agamen, 2. Aufl. Natur und Tier Verlag (Münster), 120 pp. - get paper here
  • Martin T, Guillemin M, Nivet-Mazerolles V, Landsmann C, Dubos J, Eudeline R, and Stroud J. 2017. The herpetofauna of central Uzbekistan. Amphibian & Reptile Conservation 11(1) [General Section]: 93–107 - get paper here
  • Mertens, R., and L. Müller. 1928. Liste der Amphibien und Reptilian Europas. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft. Frankfurt am Main. 41: 1-62.
  • Meyer, F.A.A. 1795. Synopsis Reptilium, novam ipsorum sistens generum methodum, nec non Gottingensium huius ordinis animalium enumerationem. Vandenhoeck und Ruprecht, Göttingen, 32 pp [for current names see Rösler & Böhme 2021] - get paper here
  • Mikaili, Peyman and Jalal Shayegh 2011. The lizards of Iran: An etymological review of families Gekkonidae, Eublepharidae, Anguidae, Agamidae. Annals of Biological Research 2 (5): 22-37
  • Milto, Konstantin D.; Andrei V. Barabanov 2012. A Catalogue of the Agamid and Chamaeleonid Types in the Collection of the Zoological Institute, Russian Academy of Sciences, St. Petersburg. Russ. J. Herpetol. 19 (2): 155-170 - get paper here
  • Mishagina J.V. 1993. Trophical connections of lizard, Phrynocephalus mystaceus, in Eastern Kara Kum. Bull. MOIP. Otd. Biol. 98 (5): 49-60 [in Russian]
  • Mishagina J.V. 2008. Ecological groups of preys in Eastern Kara Kum psammobiont lizards, Eremias (Lacertidae) and Phrynocephalus (Agamidae). The Problems of Herpetology (Proceedings of the 3rd Meeting of the Nikolsky Herpetological Society). Saint-Petersburg, 2008, pp. 298-307 [in Russian] - get paper here
  • Molavi, F.; T. Mohammadian-kalat; B.H. Kiabi; H.G. Kami; M. Yazdanpanahi 2014. New distribution Record of Phrynocephalus mystaceus Pallas 1776 (Sauria: Agamidae) from Iran. Check List 10 (2):450-452 - get paper here
  • Müller, Friedrich 1881. Zweiter Nachtrag zum Katalog der herpetologischen Sammlung des Basler Museums [1881 printed on paper]. Verh. naturf. Ges. Basel 7 (2): 166-174. - get paper here
  • Neymark, L. A. and B. D. Vasiliev 2016. ECOLOGY OF SEMIDESERT LIZARDS (REPTILIA, SQUMATA) UNDER PLANT COLONIZATION OF SANDS IN THE LOWER REACHES OF THE VOLGA RIVER (in Russian). ЗООЛОГИЧЕСКИЙ ЖУРНАЛ 95 (6): 720–727 - get paper here
  • Okshtein I., Cherlin V. 2021. Unusual evening activity of toad-headed agamas (Phrynocephalus mystaceus) on the Sarykum dune (Republic of Dagestan, Russian Federation) [in Russian] Principy èkologii 11 (4): 3‒2
  • Pallas, P.S. 1776. Reise durch verschiedene Provinzen des russischen Reichs. Kais. Akad. Wiss., St. Petersburg, Vol. 3: 703 - get paper here
  • Pang, Junfeng; Yuezhao Wang, Yang Zhong, A. Rus Hoelzel, Theodore J. Papenfuss, Xiaomao Zeng, Natalia B. Ananjeva and Ya-ping Zhang 2003. A phylogeny of Chinese species in the genus Phrynocephalus (Agamidae) inferred from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution 27: 398-409 - get paper here
  • Polynova G. V. 2016. An account of the activity of the secret toadhead agama (Phrynocephalus mystaceus mystaceus Pall. 1776) during assessments of its population. PROCEEDINGS OF THE 6th CONGRESS OF THE A. M. NIKOLSKY HERPETOLOGICAL SOCIETY pp. 120-125 - get paper here
  • Polynova G. V., Polynova O. E. 2023. Foster’s or island rule in populations of Phrynocephalus mystaceus and Eremias velox (Reptilia, Lacertilia) on the sandy massif Sarykum. Current Studies in Herpetology 23 (3): 154 - get paper here
  • Raimbekova, Eugenia 2011. The Communicative Behavior in Phrynocephalus mystaceus. HerpNation 2.4 (7): 40-47 - get paper here
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Roitberg, E. S., Mazanaeva, L.F., Ilyina, E. & Orlova, V.F. 2000. Die Echsen D<br>agestans (Nordkaukasus, Russland): Artenliste und aktuelle Verbreitungsdaten (Reptilia: Sauria: Gekkonidae, Agamidae, Anguidae, Scincidae et Lacertidae). Faunist. Abh. Staatl. Mus. Tierk. Dresden 22 (8) - get paper here
  • Rösler, Herbert & Wolfgang Böhme 2021. Das Leben von Friedrich Albrecht Anton Meyer (1768 – 1795) und die Herpetologie in seinen Schriften, unter besonderer Berücksichtigung der „Synopsis Reptilium“. Sekretär 21: 83-129
  • Semenov, D.V. and Shenbrot, G.I. 1990. Species of the genus Phrynocephalus of the USSR fauna. Description of a new subspecies with remarks on the taxonomic status of Phrynocephalus mystaceus (Reptilia, Agamidae) (in Russian, English summary). Zool. Zh. (USSR) 69(5):76-81
  • Shaw, G. 1802. General Zoology, or Systematic Natural History. Vol.3, part 2. G. Kearsley, Thomas Davison, London: 313-615 - get paper here
  • Shaw, George, Nodder, Elizabeth (Eds.) 1804. The Naturalist's Miscellany [...], Vol. XV. London, Nodder & Co., plates 589-636, 177 unnumbered pages [published in monthly issues between August 1, 1803, and July 1, 1804] - get paper here
  • Shestopal A. A., Shcherbina A. A. 2024. Distribution of reptiles in Northwestern Turkmenistan. [in Russian] Current Studies in Herpetology 24 (1-2): 102 - get paper here
  • Shestopal A., Amanov A., Ovezov T. 2021. An overview of reptiles from the Kaplankyr Nature Reserve and adjacent territories. [in Russian] Proc. Inst. Zool., Kazakhstan, pp. 105-126 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • ŠMÍD, JIŘÍ; JIŘÍ MORAVEC, PETR KODYM, LUKÁŠ KRATOCHVÍL, SEYYED SAEED HOSSEINIAN YOUSEFKHANI, ESKANDAR RASTEGAR-POUYANI & DANIEL FRYNTA 2014. Annotated checklist and distribution of the lizards of Iran. Zootaxa 3855 (1): 001–097 - get paper here
  • Solovyeva EN, Dunayev EN, Nazarov RA, Radjabizadeh M, Poyarkov Jr NA 2018. Molecular and morphological differentiation of Secret Toad-headed agama, Phrynocephalus mystaceus, with the description of a new subspecies from Iran (Reptilia, Agamidae). ZooKeys 748: 97-129 - get paper here
  • Solovyeva, E. N., Dunayev, E. A., Nazarov, R. A., Bondarenko, D. A., & Poyarkov, N. A. 2023. COI-Barcoding and Species Delimitation Assessment of Toad-Headed Agamas of the Genus Phrynocephalus (Agamidae, Squamata) Reveal Unrecognized Diversity in Central Eurasia. Diversity 2023, 15, x - get paper here
  • Szczerbak, N.N. 2003. Guide to the Reptiles of the Eastern Palearctic. Krieger, Malabar, FL, 260 pp. - get paper here
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • WHITING, MARTIN J.; DANIEL W.A. NOBLE & YIN QI. 2022. A potential deimatic display revealed in a lizard. Biological Journal of the Linnean Society 136(3): 455–465. - get paper here
  • Zhao,E. & Adler,K. 1993. Herpetology of China. SSAR, Oxford/Ohio, 1-522
  • Zheng, P., Liang, T., An, J., & Shi, L. 2020. Morphological function of toe fringe in the sand lizard Phrynocephalus mystaceus. Scientific Reports, 10(1): 1-10 - get paper here
 
External links  
Is it interesting? Share with others:

As link to this species use URL address:

https://reptile-database.reptarium.cz/species?genus=Phrynocephalus&species=mystaceus

without field 'search_param'. Field 'search_param' is used for browsing search result.



Please submit feedback about this entry to the curator