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Phrynocephalus mystaceus (PALLAS, 1776)

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Higher TaxaAgamidae (Agaminae), Sauria (lizards) 
SubspeciesPhrynocephalus mystaceus mystaceus (PALLAS 1776)
Phrynocephalus mystaceus galli KRASSOWSKY 1932 
Common NamesE: Secret Toadhead Agama
G: Bärtiger Krötenkopf 
SynonymLacerta mystacea PALLAS 1776: 702
Lacerta aurita — SHAW & NODDER 1803: plate 594
Phrynocephalus mystacia — KAUP 1827
Phrynocephalus mystaceus — BOULENGER 1885: 379
Phrynocephalus mystaceus mystaceus natio dagestanica KRASSOWSKY 1932
Phrynocephalus mystaceus galli KRASSOWSKY 1932
Megalochilus mystaceus — ANANJEVA 1986
Phrynocephalus mystaceus — WERMUTH 1967: 84
Phrynocephalus mystaceus — ENGELMANN et al 1993

Phrynocephalus mystaceus mystaceus (PALLAS 1776)
Lacerta aurita GMELIN 1789: 1073 (nom. subst. pro Lacerta mystacea)
Gekko auritus — LATREILLE 1801: 61.
Lacerta lobata SHAW 1802: 244 (nom. subst. pro Lacerta aurita)
Agama aurita — DAUDIN 1802: 429
Agama mystacea — MERREM 1820: 53
Megalochilus auritus — EICHWALD 1831: 185
Phrynocephalus auritus — EVERSMANN 1834: 360
Phrynocephalus auritus — DUMÉRIL & BIBRON 1837: 524
Saccostoma auritum — FITZINGER 1843: 87
Phrynocephalus mystaceus — BOETTGER 1880: 260
Phrynocephalus mystaceus mystaceus — KRASSOWSKY 1932: 227
Phrynocephalus mystaceus mystaceus — WERMUTH 1967: 84
Phrynocephalus mystaceus mystaceus — MANTHEY & SCHUSTER 1999: 90
Phrynocephalus mystaceus mystaceus — BARABANOV & ANANJEVA 2007

Phrynocephalus mystaceus galli KRASSOWSKY 1932
Phrynocephalus mystaceus Galli KRASSOWSKY 1932: 227
Phrynocephalus mystaceus aurantiacaudatus SEMENOV & SHENBROT 1990: 79
Phrynocephalus mystaceus galli — BARABANOV & ANANJEVA 2007
Phrynocephalus mystaceus aurantiacaudatus — SINDACO & JEREMČENKO 2008
Phrynocephalus mystaceus galli — SINDACO & JEREMČENKO 2008
Phrynocephalus mystaceus galli — MILTO & BARABANOV 2012 
DistributionSoviet Union, NW cost of Caspian Sea to E Kazakhstan, Russia (Dagestan, Astrakhan Oblast), Kazakhstan, Turkmenistan, Tajikistan, Uzbekistan, Kyrgyzstan, NE Iran, Afghanistan, NW China (W Xinjiang)

aurantiacaudatus: Kazakhstan
galli: Type locality: Repeteka, Turkestan.

galli: Turkmenistan, Uzbekistan, Kazakhstan, northeastern and eastern Iran and adjacent Afghanistan and Xinjiang, China; Terra typica: “Aus Repeteka andetrifft” [= vicinity of Repetek station, approximately 38º35' N, 63º11' E, Lebapsky Region, Turkmenistan].

Type locality: East Caucasus. Restricted to “Naryn-Steppe, an der Nordküste des kaspischen Maeeres” by MERTENS & MÜLLER 1928; Restricted by neotype designation to Ryn Peski (Ryn Sands), Uralskaya Region, north-western Kazakh-
stan, by BARABANOV & ANANJEVA 2007. Map legend:
Type locality - Type locality.
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
TypesNeotype: ZISP 8735.1, adult male, designated by BARABANOV & ANAJEVA 2007 
CommentSynonymy and subspecies mainly after KHALIKOV & ANANJEVA (pers. comm.), WERMUTH 1967, and BARABANOV & ANANJEVA 2007. SEMENOV & SHENBROT 1990 considered galli as a synonym of P. mystaceus.

Reproduction: Oviparous.

Size: Largest species within genus (up to 25 cm total length). 
References
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  • Ananjeva, N. B. 1986. O rodovoj samostoyatelnosti ushastoj kriglogolovki, Megalochilus mystaceus (PALLAS 1776). [On the validity of Megalochilus mystaceus (PALLAS 1776).]. Trud. Zool. Inst. AN SSSR, Leningrad 157: 4-13
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