You are here » home advanced search Phrynocephalus mystaceus

Phrynocephalus mystaceus (PALLAS, 1776)

Can you confirm these amateur observations of Phrynocephalus mystaceus?

Add your own observation of
Phrynocephalus mystaceus »

Find more photos by Google images search: Google images

Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards) 
SubspeciesPhrynocephalus mystaceus mystaceus (PALLAS 1776)
Phrynocephalus mystaceus galli KRASSOWSKY 1932 
Common NamesE: Secret Toadhead Agama
G: Bärtiger Krötenkopf 
SynonymLacerta mystacea PALLAS 1776: 702
Lacerta aurita — SHAW & NODDER 1803: plate 594
Phrynocephalus mystacia — KAUP 1827
Phrynocephalus mystaceus — BOULENGER 1885: 379
Phrynocephalus mystaceus mystaceus natio dagestanica KRASSOWSKY 1932
Phrynocephalus mystaceus galli KRASSOWSKY 1932
Megalochilus mystaceus — ANANJEVA 1986
Phrynocephalus mystaceus — WERMUTH 1967: 84
Phrynocephalus mystaceus — ENGELMANN et al 1993
Phrynocephalus mystaceus — ŠMÍD et al. 2014

Phrynocephalus mystaceus mystaceus (PALLAS 1776)
Lacerta aurita GMELIN 1789: 1073 (nom. subst. pro Lacerta mystacea)
Gekko auritus — LATREILLE 1801: 61.
Lacerta lobata SHAW 1802: 244 (nom. subst. pro Lacerta aurita)
Agama aurita — DAUDIN 1802: 429
Agama mystacea — MERREM 1820: 53
Agama aurita — LIECHTENSTEIN 1823: 142
Megalochilus auritus — EICHWALD 1831: 185
Phrynocephalus auritus — EVERSMANN 1834: 360
Phrynocephalus auritus — DUMÉRIL & BIBRON 1837: 524
Saccostoma auritum — FITZINGER 1843: 87
Phrynocephalus mystaceus — BOETTGER 1880: 260
Phrynocephalus mystaceus mystaceus — KRASSOWSKY 1932: 227
Phrynocephalus mystaceus mystaceus — WERMUTH 1967: 84
Phrynocephalus mystaceus mystaceus — MANTHEY & SCHUSTER 1999: 90
Phrynocephalus (Phrynocephalus) mystaceus mystaceus — BARABANOV & ANANJEVA 2007

Phrynocephalus mystaceus galli KRASSOWSKY 1932
Phrynocephalus mystaceus Galli KRASSOWSKY 1932: 227
Phrynocephalus mystaceus aurantiacaudatus SEMENOV & SHENBROT 1990: 79
Phrynocephalus (Phrynocephalus) mystaceus galli — BARABANOV & ANANJEVA 2007
Phrynocephalus mystaceus aurantiacaudatus — SINDACO & JEREMČENKO 2008
Phrynocephalus mystaceus galli — SINDACO & JEREMČENKO 2008
Phrynocephalus mystaceus galli — MILTO & BARABANOV 2012 
DistributionSoviet Union, NW cost of Caspian Sea to E Kazakhstan, Russia (Dagestan, Astrakhan Oblast), Kazakhstan, Turkmenistan, Tajikistan, Uzbekistan, Kyrgyzstan, NE Iran, Afghanistan, NW China (W Xinjiang)

aurantiacaudatus: Kazakhstan
galli: Type locality: Repeteka, Turkestan.

galli: Turkmenistan, Uzbekistan, Kazakhstan, northeastern and eastern Iran and adjacent Afghanistan and Xinjiang, China; Terra typica: “Aus Repeteka andetrifft” [= vicinity of Repetek station, approximately 38º35' N, 63º11' E, Lebapsky Region, Turkmenistan].

Type locality: East Caucasus. Restricted to “Naryn-Steppe, an der Nordküste des kaspischen Maeeres” by MERTENS & MÜLLER 1928; Restricted by neotype designation to Ryn Peski (Ryn Sands), Uralskaya Region, north-western Kazakh-
stan, by BARABANOV & ANANJEVA 2007. Map legend:
Type locality - Type locality.
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
TypesNeotype: ZISP 8735.1, adult male, designated by BARABANOV & ANAJEVA 2007 
CommentSynonymy and subspecies mainly after KHALIKOV & ANANJEVA (pers. comm.), WERMUTH 1967, and BARABANOV & ANANJEVA 2007. SEMENOV & SHENBROT 1990 considered galli as a synonym of P. mystaceus.

Distribution: See map in SMID et al. 2014 for distribution in Iran. 
  • Ananjeva, N. & Orlov, N. 2005. Lizards of North Eurasia. Reptilia (GB) (38): 54-63 - get paper here
  • Ananjeva, N. B. 1986. O rodovoj samostoyatelnosti ushastoj kriglogolovki, Megalochilus mystaceus (PALLAS 1776). [On the validity of Megalochilus mystaceus (PALLAS 1776).]. Trud. Zool. Inst. AN SSSR, Leningrad 157: 4-13
  • Ananjeva, Natalia B. 2012. Research on Asian Agamid Lizards: Contribution of Zoological Institute, Russian Academy of Sciences, St. Petersburg. Russ. J. Herpetol. 19 (2): 85-100 - get paper here
  • ANANJEVA, Natalia B.; Xianguang GUO and Yuezhao WANG 2011. Taxonomic Diversity of Agamid Lizards (Reptilia, Sauria, Acrodonta, Agamidae) from China: A Comparative Analysis. Asian Herpetological Research 2 (3): 117-128 - get paper here
  • Ananjeva, Natalia B.;Tuniyev, Boris S. 1992. Historical biogeography of the Phrynocephalus species of the USSR. Asiatic Herpetological Research 4: 76-98 - get paper here
  • Ananjeva, Natalija B. 1981. Phrynocephalus mystaceus - Bärtiger Krötenkopf. In: Böhme,W. (ed.), Handbuch der Reptilien und Amphibien Europas, Band I., Echsen I. Akademische Verlagsgesellschaft, Wiesbaden, pp. 203-216
  • Ananjeva, NB, Orlov NL; Khalikov RG; Darevsky IS; Ryabov IS & Barabanov AV 2006. The Reptiles of North Eurasia. Taxonomic Diversity, Distribution, Conservation Status [this comprises the territory of the former Soviet Union and Mongolia]. Pensoft Series Faunistica 47, 250 pp.
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • BARABANOV, A.V. & N.B. ANANJEVA 2007. Catalogue of the available scientific species-group names for lizards of the genus Phrynocephalus Kaup, 1825 (Reptilia, Sauria, Agamidae). Zootaxa 1399: 1-56 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Davletbakov, A. T.; L. A. Kustareva, D. A. Milko, A. N. Ostastshenko, S. S. Sagymbaev, S. A. Toropov, N. V. Trotchenko 2015. Genetical Fund Cadastre of Kyrgyz Republic. Vol. IV. Phylum Chordata (chordates). Bishkek, 128 p. + XXXVI color tables - get paper here
  • Dieckmann, M. 2011. Bemerkungen zur Haltung des Bärtigen Krötenkopf Phrynocephalus mystaceus (PALLAS 1776). Iguana Rundschreiben 24 (1): 21-26
  • Dujsebayeva, T. N. (ed.) 2010. Short review of last changes in the checklist of amphibians and reptiles of Kazakhstan. [in Russian]. In: Dujsebayeva, T. N. (ed.) Herpetological Researches in Kazakhstan and adjacent countries, p. 37-52
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Engelmann,W.E. et al. 1993. Lurche und Kriechtiere Europas. Neumann Verlag (Radebeul, Germany), 440 pp.
  • Eversmann,E. 1834. Lacertae Imperii Rossici variis in itineribus meis observatae. [Lacerta sylvicola]. Nouv. Mém. Soc. imp. Natur. (Moscow) 3: 339-358 - get paper here
  • Gmelin, J.F. 1789. Caroli a Linné Systema naturae. 13. ed., Tom 1 Pars 3. G. E. Beer, Lipsiae. 1033-1516 pp. - get paper here
  • Hoogmoed, M.S. 1980. Herpetologische waarnemingen in Ghana [part 3]. Lacerta 38 (5): 42-48 - get paper here
  • Krassowsky, D.B. 1932. Beitrag zur Systematik von Phrynocephalus mystaceus. Zool. Anz. 97: 225-228
  • Kukushkin O.V., Doronin I.V., Tuniyev B.S., Ananjeva N.B., Doronina M.A. 2017. Introduction of Amphibians and Reptiles at the Caucasus and the Crimea: an Overview and Some Actual Data. (in Russian) Current Studies in Herpetology 17 (3-4): 157–197 . DOI: 10.18500/1814-6090-2017-17-3-4-157-197 - get paper here
  • Lambert, M.R.K. 2002. Preliminary observations on herpetofaunal diversity in the Almaty region, Southern Kazakhstan (September 1998). Herpetological Bulletin (79): 7-13 - get paper here
  • Lichtenstein 1823. Amphibien. in: Evermann, Reise nach Buchara: 144. - get paper here
  • Macey, J. R., J. A. Schulte II, A. Larson, N. B. Ananjeva, Y. Wang, R. Pethiyagoda, N. Rastegar-Pouyani, T. J. Papenfuss 2000. Evaluating trans-Tethys migration: an example using acrodont lizard phylogenetics. Systematic Biology 49 (2): 233-256 - get paper here
  • Manthey,U. & SCHUSTER,N. 1999. Agamen, 2. Aufl. Natur und Tier Verlag (Münster), 120 pp. - get paper here
  • Martin T, Guillemin M, Nivet-Mazerolles V, Landsmann C, Dubos J, Eudeline R, and Stroud J. 2017. The herpetofauna of central Uzbekistan. Amphibian & Reptile Conservation 11(1) [General Section]: 93–107 - get paper here
  • Mertens, R., and L. Müller. 1928. Liste der Amphibien und Reptilian Europas. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft. Frankfurt am Main. 41: 1-62.
  • Milto, Konstantin D.; Andrei V. Barabanov 2012. A Catalogue of the Agamid and Chamaeleonid Types in the Collection of the Zoological Institute, Russian Academy of Sciences, St. Petersburg. Russ. J. Herpetol. 19 (2): 155-170 - get paper here
  • Mishagina J.V. 1993. Trophical connections of lizard, Phrynocephalus mystaceus, in Eastern Kara Kum. Bull. MOIP. Otd. Biol. 98 (5): 49-60 [in Russian]
  • Mishagina J.V. 2008. Ecological groups of preys in Eastern Kara Kum psammobiont lizards, Eremias (Lacertidae) and Phrynocephalus (Agamidae). The Problems of Herpetology (Proceedings of the 3rd Meeting of the Nikolsky Herpetological Society). Saint-Petersburg, 2008, pp. 298-307 [in Russian] - get paper here
  • Pallas, P.S. 1776. Reise durch verschiedene Provinzen des russischen Reichs. Kais. Akad. Wiss., St. Petersburg, Vol. 3: 703 - get paper here
  • Pang, Junfeng; Yuezhao Wang, Yang Zhong, A. Rus Hoelzel, Theodore J. Papenfuss, Xiaomao Zeng, Natalia B. Ananjeva and Ya-ping Zhang 2003. A phylogeny of Chinese species in the genus Phrynocephalus (Agamidae) inferred from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution 27: 398-409 - get paper here
  • Polynova G. V. 2016. An account of the activity of the secret toadhead agama (Phrynocephalus mystaceus mystaceus Pall. 1776) during assessments of its population PROCEEDINGS OF THE 6th CONGRESS OF THE A. M. NIKOLSKY HERPETOLOGICAL SOCIETY pp. 120-125 - get paper here
  • Raimbekova, Eugenia 2011. The Communicative Behavior in Phrynocephalus mystaceus. HerpNation 2.4 (7): 40-47 - get paper here
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Roitberg, E. S., Mazanaeva, L.F., Ilyina, E. & Orlova, V.F. 2000. Die Echsen Dagestans (Nordkaukasus, Russland): Artenliste und aktuelle Verbreitungsdaten (Reptilia: Sauria: Gekkonidae, Agamidae, Anguidae, Scincidae et Lacertidae). Faunist. Abh. Staatl. Mus. Tierk. Dresden 22 (8) - get paper here
  • Semenov, D.V. and Shenbrot, G.I. 1990. Species of the genus Phrynocephalus of the USSR fauna. Description of a new subspecies with remarks on the taxonomic status of Phrynocephalus mystaceus (Reptilia, Agamidae) (in Russian, English summary). Zool. Zh. (USSR) 69(5):76-81
  • Shaw, G. 1802. General Zoology, or Systematic Natural History. Vol.3, part 2. G. Kearsley, Thomas Davison, London: 313-615 - get paper here
  • Shaw, George, Nodder, Elizabeth (Eds.) 1804. The Naturalist's Miscellany [...], Vol. XV. London, Nodder & Co., plates 589-636, 177 unnumbered pages [published in monthly issues between August 1, 1803, and July 1, 1804] - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • ŠMÍD, JIŘÍ; JIŘÍ MORAVEC, PETR KODYM, LUKÁŠ KRATOCHVÍL, SEYYED SAEED HOSSEINIAN YOUSEFKHANI, ESKANDAR RASTEGAR-POUYANI & DANIEL FRYNTA 2014. Annotated checklist and distribution of the lizards of Iran. Zootaxa 3855 (1): 001–097
  • Szczerbak, N.N. 2003. Guide to the Reptiles of the Eastern Palearctic. Krieger, Malabar, FL, 260 pp. - get paper here
  • Zhao,E. & Adler,K. 1993. Herpetology of China. SSAR, Oxford/Ohio, 1-522
External links  
Is it interesting? Share with others:

Please submit feedback about this entry to the curator