Phrynocephalus putjatai BEDRIAGA, 1909
Can you confirm these amateur observations of Phrynocephalus putjatai?
|Higher Taxa||Agamidae (Agaminae), Sauria, Iguania, Squamata (lizards)|
|Common Names||Chinese: 贵德沙蜥|
|Synonym||Phrynocephalus putjatai BEDRIAGA 1909: 302|
Phrynocephalus putiatae — TZAREVSKY 1929: 419 (unjustified emendation).
Phrynocephalus putjatai — WERMUTH 1967: 86
Phrynocephalus putjaitai — ZHAO 1997: 31 (unjustified emendation)
Phrynocephalus putjatia — WANG et al., 2002: 372 (unjustified emendation).
Phrynocephalus putjatia — GUO & WANG 2007
Phrynocephalus (Oreosaura) putjatai — BARABANOV & ANANJEVA 2007
Phrynocephalus putjatae — JI et al. 2009
Phrynocephalus guinanensis JI, WANG & WANG 2009
Phrynocephalus putjatia — JIN et al. 2013
Phrynocephalus putjatia — WANG et al. 2017
Phrynocephalus putjatia — JIN et al. 2018
Phrynocephalus guinanensis — JIN et al. 2018
Phyrnocephalus guinanensis — JIN & BROWN 2019 (in error)
Phrynocephalus guinanensis — JIN & BROWN 2019
Phrynocephalus putjatia — JIN & BROWN 2019
|Distribution||China (Guide, Qinghai Province)|
Type locality: “Gui-dui am Hoang-ho” [= Guide County (36º00' N, 101º 40' E), southeast of Qinghai (Kuku-Nor) Lake, Qinghai Province, China].
guinanensis: China (Guinan, Qinghai); Type locality: Senduo (35°34’N, 101°05’E; elevation 3350 m ASL), Guinan County, Qinghai Province, China.
|Types||Lectotype: ZISP 7447.1 (designated by Peters, 1984: 44).|
Holotype: NNU (Nanjing Normal University) P 2004.006, adult male (Fig. 1A, C); collected by Xiang Ji on 17 May 2004 [guinanensis]
|Diagnosis||Diagnosis (guinanensis): A large viviparous species of Phrynocephalus, distinguishable from congeners using the following combination of characters: up to 84 mm SVL; dorsal ground color of head, neck, trunk, limbs and tail brown with weak light brown mottling; lateral ground color of head, neck, trunk and tail light black with weak white-gray mottling in adult males, and green with weak white-gray mottling in adult females; ventral ground color of tail white-gray to black in the distal part of the tail in adult males, and totally white-gray in adult females; ventral surfaces of hind-limbs white-gray; ventral surfaces of fore-limbs brick-red in adult males, and white-gray in adult females; ventral ground color of trunk and head black in the center but, in the periphery, brick-red in adult males and white-gray in adult females, and totally white-gray in adult females; 5− 8 nasal scales; 5−8 internasal scales; 8−12 scales around parietal eye; 13−18 supraocular scales; 126−164 dorsal scales; 101−136 ventral scales; 144−186 scales around mid-body; 21−31 gular folds; 15−19 subdigital lamellae of the 4th finger; 22−29 subdigital lamellae of the 4th toe.|
|Comment||Synonymy: Bayesian species delimitation analysis did not support division of P. putjatia and P. guinanensis into separate species (JIN et al. 2013). JIN et al. 2018 confirmed their close relationship. However, Jin & Brown 2019 recognize guinanensis as a separate species again. However, despite some genetic analysis their study does not attempt to show the phylogenetic relationship between different populations of these species. The original description of P. guinanensis also remains a bit dubious and seems to be primarily based on size and color variation (only guinanensis shown, no other species).|
Phrynocephalus putjatae has been considered as a synonym of P. vlangalii. P. vlangalii and P. putjatae hybridize in the Qinghai region (JIN & LIU 2008). Of uncertain status fide BARABANOV & ANANJEVA 2007 but considered as a valid species by GUO et al. 2012.
Habitat (guinanensis): sand dunes with very sparse vegetation.
|Etymology||Named after Mr. D.W. Putjata, “für seine amphibologischen Zuwendungen an das Museum”.|
The specific epithet of P. guinanensis is a latinization of the name Guinan County, Qinghai Province, China.