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Phrynocephalus raddei BOETTGER, 1888

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards) 
SubspeciesPhrynocephalus raddei boettgeri VON BEDRIAGA 1906
Phrynocephalus raddei raddei BOETTGER 1888 
Common Names 
SynonymPhrynocephalus raddei BOETTGER 1888: 262
Phrynocephalus reticulatus EICHWALD 1831 (fide KHALIKOV, pers. comm.)
Phrynocephalus raddei var. bilkewitschi NIKOLSKY 1915: 194
Phrynocephalus reticulatus boettgeri — LANGERWERF 1980
Phrynocephalus raddei — ANDERSON 1999
Phrynocephalus raddei — MACEY et al. 2000
Phrynocephalus (Phrynocephalus) raddei — BARABANOV & ANANJEVA 2007
Phrynocephalus raddei — SINDACO & JEREMČENKO 2008
Phrynocephalus raddei raddei BOETTGER 1888

Phrynocephalus raddei boettgeri VON BEDRIAGA 1906
Phrynocephalus raddei var. boettgeri BEDRIAGA In NIKOLSKY 1905: 468
Phrynocephalus raddei var. zardunyi NIKOLSKY 1915: 194
Phrynocephalus raddei boettgeri — SINDACO & JEREMČENKO 2008
Phrynocephalus raddei boettgeri — MILTO & BARABANOV 2012
Phrynocephalus raddei boettgeri — ANANJEVA et al. 2020 
DistributionS Turkmenia, S Uzbekistan, SW Tadjikistan, Iran, N Afghanistan

Type locality: "Perewalnaja an der transcaspischen Bahn" [= Perevalnaja railroad station, southwestern Turkmenistan].

zarudnyi: Type locality: “Kelif, Buchara orient.” [Kelif, Lebap Province, Turkmenistan, 37.36 N 66.31 E].  
TypesLectotype: SMF 10247 (designated by Mertens, 1967: 63), (formerly 4369a), syntype: NMW 23344
Lectotype: ZISP 6117.1, adult female, (designated by Barabnov & Ananjeva 2007: 15) [boettgeri]
Lectotype: ZISP 10722.1, adult male (designated by Barabanov & Ananjeva, 2007) [zardunyi] 
DiagnosisDiagnosis: Upper surface of snout fairly steeply truncated, nostrils not visible or faintly noticeable when viewed from above; no cutaneous fold at angle of mouth; dorsal scales subequal, homogeneous; no transverse fold of skin across back of neck; sides ofhead and neck without projecting fringe-like scales; no fringe of scales on posterior margin of thigh and sides of base of tail; no enlarged scales along flank; no tubercular scales on dorsum; scales of upper surfaces of limbs and midline of back prominently keeled. A half-moon-shaped red patch on each scapular region, 21-24 lamellae under fourth toe; ridges on subdigital lamellae of third toe approximately uniformly developed in 3 rows (Anderson 1999: 96). 
CommentSynonymy: Listed as synonym of Phrynocephalus reticulatus by WERMUTH 1967. P. r. boettgeri is a synonym of raddei according to BARABANOV & ANANJEVA 2007. 
  • Ananjeva, Natalia B.;Tuniyev, Boris S. 1992. Historical biogeography of the Phrynocephalus species of the USSR. Asiatic Herpetological Research 4: 76-98 - get paper here
  • Ananjeva, NB, Orlov NL; Khalikov RG; Darevsky IS; Ryabov IS & Barabanov AV 2006. The Reptiles of North Eurasia. Taxonomic Diversity, Distribution, Conservation Status [this comprises the territory of the former Soviet Union and Mongolia]. Pensoft Series Faunistica 47, 250 pp.
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • BARABANOV, A.V. & N.B. ANANJEVA 2007. Catalogue of the available scientific species-group names for lizards of the genus Phrynocephalus Kaup, 1825 (Reptilia, Sauria, Agamidae). Zootaxa 1399: 1-56 - get paper here
  • Bedriaga, JACQUES VON 1907. Neue Saurier aus Russisch-Asien. Annu. Mus. Zool. Acad. Impér. Sci. St.-Petersbourg, 10 [1905]: 210-243 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boettger, O. 1888. Über die Reptilien und Batrachier Transcaspiens. Zool. Anz. 11: 259-263 - get paper here
  • Joger,U. 1991. A molecular phylogeny of agamid lizards. Copeia 1991 (3): 616-622 - get paper here
  • Langerwerf, B. 1980. De herpetofauna van Oezbekistan (3). Lacerta 38 (4): 37-40 - get paper here
  • Macey, J. R., J. A. Schulte II, A. Larson, N. B. Ananjeva, Y. Wang, R. Pethiyagoda, N. Rastegar-Pouyani, T. J. Papenfuss 2000. Evaluating trans-Tethys migration: an example using acrodont lizard phylogenetics. Systematic Biology 49 (2): 233-256 - get paper here
  • Milto, Konstantin D.; Andrei V. Barabanov 2012. A Catalogue of the Agamid and Chamaeleonid Types in the Collection of the Zoological Institute, Russian Academy of Sciences, St. Petersburg. Russ. J. Herpetol. 19 (2): 155-170 - get paper here
  • Nikolsky, A. M. 1905. Herpetologia rossica. Mem. Acad. Imp. Sci., Phys.-Math. Classe, St. Pétersbourg, XVII (1): 1-518
  • Nikolsky, A. M. 1915. Faune de la Russie, Reptiles Vol. 1 Chelonia et Sauria(Reptiles of Russia and adjacent countries. Reptiles (Reptilia) Vol. 1) [in Russian]. Imper. Acad. Sci., Petrograd, 532 pp. [Translated from Russian by the Israel Program for Scientific Translations 1963]
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Szczerbak, N.N. 2003. Guide to the Reptiles of the Eastern Palearctic. Krieger, Malabar, FL, 260 pp. - get paper here
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