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Phrynonax shropshirei BARBOUR & AMARAL, 1924

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Shropshire's Puffing Snake 
SynonymPhrynonax shropshirei BARBOUR & AMARAL 1924: 131
Pseustes poecilonotus shropshirei — AMARAL 1929: 317
Pseustes shropshirei — BRONGERSMA 1937: 6
Pseustes shropshirei — PETERS & OREJAS-MIRANDA 1970: 259
Pseustes shropshirei — KORNACKER 1999: 129
Pseustes poecilonotus shropshirei — PORRAS & SOLÓRZANO 2006
Pseustes shropshirei — MORENO-ARIAS et al. 2008
Phrynonax shropshirei — JADIN et al. 2013
Pseustes shropshirei — WALLACH et al. 2014: 612 
DistributionCosta Rica, Panama, Pacific Colombia, Venezuela, Ecuador, (Barinas [HR 34: 389] etc, KORNACKER 1999), Peru

elevation 0 -1300 m

Type locality: Fort Sherman, Canal Zone, Panama.  
Reproductionoviparous 
TypesHolotype: MCZ 18819 
DiagnosisAdditional details, e.g. a detailed description or comparisons (1810 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSynonymy: This species may be synonymous to P. poecilonotus as the latter is extremely variable. Pseustes shropshirei (Barbour and Amaral 1924) appears to be based on a typical obscurely banded and light-spotted example of Panamanian P. poecilonotus, as pointed out by Amaral (1929). See SAVAGE 2002 for a discussion. Note that the deadline for WALLACH et al. 2014 was late 2012, hence Jadin et al. 2013 is the most recent analysis of this genus.

Distribution: This species has been reported from Peru, but without specific locality (T. Doan, pers. comm. 30 Apr 2012). Not in Venezuela fide NATERA-MUMAW et al. 2015, but there fide Señaris et al. 2018.

Habitat: partly arboreal (Harrington et al. 2018). 
EtymologyNamed after James B. Shropshire, “...Chief Sanitary Inspector, U.S. Army, Canal Zone...”. 
References
  • Almendariz, A. 1991. Anfibios y Reptiles [del Ecuador]. Rev. Politecnica. XVI (3): 89-162.
  • Barbour, Thomas & Amaral,A. 1924. Notes on some Central American snakes. Occasional Papers of the Boston Society of Natural History 5: 129-132 - get paper here
  • Brongersma, L. D. 1937. Herpetological Notes XIV - XVI. XV. The type of Dipsas dieperinkii Schlegel, 1837, and the genus Pseustes Fitzinger, 1843. Zoologische Mededelingen 20: 1-10 - get paper here
  • Carvajal-Cogollo, J.E.; L.E. Rojas-Murcia. & G. Cárdenas-Arévalo 2020. Reptiles del Caribe colombiano/ Reptiles of the Colombian Caribbean. Tunja: Editorial UPTC, 268 pp. - get paper here
  • Castro-Herrera, F. & Vargas-Salinas, F. 2008. Anfibios y reptiles en el departamento del Valle del Cauca, Colombia. Biota Colombiana 9 (2): 251 - 277 - get paper here
  • Freiberg, M.A. 1982. Snakes of South America. T.F.H. Publications, Neptune City, NJ, 189 pp. - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Jadin, R. C., Burbrink, F. T., Rivas, G. A., Vitt, L. J., Barrio-Amorós, C. L. and Guralnick, R. P. 2013. Finding arboreal snakes in an evolutionary tree: phylogenetic placement and systematic revision of the Neotropical birdsnakes. Journal of Zoological Systematics and Evolutionary Research. 52 (3): 257–264; doi: 10.1111/jzs.12055 - get paper here
  • Kornacker,P.M. 1999. Checklist and key to the snakes of Venezuela. PaKo-Verlag, Rheinbach, Germany, 270 pp.
  • Lehr, E. 2002. Amphibien und Reptilien in Peru. Natur und Tier-Verlag (Münster), 208 pp. - get paper here
  • Moreno-Arias, R.A., G.F. Medina-Rangel & O.V. Castaño-Mora 2008. Lowland reptiles of Yacopí (Cundinamarca, Colombia). Rev. Acad. Colomb. Cienc. 32 (122): 93-103 - get paper here
  • Páez, V.P. et al. 2002. Guía de campo de algunas especies de anfibios y reptiles de Antioquia. Universidad de Antioquia, - get paper here
  • Pérez-Santos,C. & Moreno, A.G. 1988. Ofidios de Colombia. Museo reegionale di Scienze Naturali, Torino, Monographie VI, 517 pp.
  • Peters , J. A. 1960. The snakes of Ecuador; check list and key. Bull. Mus. Comp. Zool. Harvard 122: 489-541 - get paper here
  • Pinto-Erazo, M.A., Calderón-Espinosa, M.L., Medina-Rangel, G.F. & Méndez-Galeano, M.A. 2020. Herpetofauna from two municipalities of southwestern Colombia. Biota Colombiana 21(1): 41-57. DOI: 10.21068/c2020.v21n01a04 - get paper here
  • Rivas Fuenmayor, G. & Molina,C.R. 1998. Geographic distribution. Pseustes shropshirei. Herpetological Review 29 (3): 178 - get paper here
  • RIVAS, GILSON A.; CÉSAR R. MOLINA, GABRIEL N. UGUETO, TITO R. BARROS, CÉSAR L. BAR- RIO-AMORÓS & PHILIPPE J. R. KOK 2012. Reptiles of Venezuela: an updated and commented checklist. Zootaxa 3211: 1–64 - get paper here
  • Rojas-Morales, Julián Andrés, Héctor Fabio Arias-Monsalve y Gustavo A. González-Durán 2014. Anibios y reptiles de la región centro-sur del departamento de Caldas, Colombia. Biota Colombiana 15 (1): 73-93 - get paper here
  • Señaris, J. Celsa; María Matilde Aristeguieta Padrón, Haidy Rojas Gil y Fernando J. M. Rojas-Runjaic 2018. Guía ilustrada de los anfibios y reptiles del valle de Caracas, Venezuela. Ediciones IVIC, Instituto Venezolano de Investigaciones Científicas (IVIC). Caracas, Venezuela. 348 pp.
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Zuluaga-Isaza, Juan Camilo, Julián Andrés Rojas-Morales, Román Felipe Díaz-Ayala and Viviana Andrea Ramírez-Castaño 2015. Pseustes shropshirei (Shropshire's puffing snake) diet. Herpetological Review 46 (4): 649. - get paper here
 
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