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Phrynosoma goodei STEJNEGER, 1893

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Higher TaxaPhrynosomatidae, Phrynosomatinae, Phrynosomatini; Iguania, Sauria, Squamata (lizards)
Common NamesE: Sonoran Horned Lizard
G: Sonora-Wüstenkrötenechse
S: Camaleón de Sonora 
SynonymPhrynosoma platyrhinos goodei STEJNEGER 1893: 191
Anota goodei — COPE 1896: 834
Anota goodei — COPE 1900
Phrynosoma platyrhinos goodei — PIANKA 1991
Phrynosoma goodei — MULCAHY et al. 2006
Phrynosoma goodei — COLLINS & TAGGART 2009
Phrynosoma (Doliosaurus) goodei — CROTHER et al. 2012
Phrynosoma platyrhinos goodei — KÖHLER 2021
Phrynosoma goodei — HEIMES 2022 
DistributionUSA (SW Arizona), Mexico (NW Sonora)

Type locality: "Coast deserts of the state of Sonora, Mexico," restricted to Puerto Libertad by SMITH & TAYLOR 1950.  
TypesHolotype: USNM 8567a 
DiagnosisDiagnosis: 3 temporal horns are enlarged, the one posterior equal in size to occipital horns; the 3 posterior chin shields are greatly enlarged and pointed (Pianka 1991). 
CommentMULCAHY et al. 2006 elevated Phrynosoma platyrhinos goodei to full species status based on mtDNA sequences. MULCAHY et al. 2006 also presented evidence for goodei x mcalli hybrids.

Group: Belongs to the Doliosaurus clade fide LEACHE & MCGUIRE 2006. 
EtymologyNamed after George Brown Goode (1851-1896), American ichthyologist, assistant to Baird, and Assistant Secretary ofthe Smithsonian from 1872. 
  • Aguilar-Morales, Cecilia and Thomas R. Van Devender 2018. Horned Lizards (Phrynosoma) of Sonora, Mexico: Distribution and Ecology. Sonoran Herpetologist 31 (3): 40-50 - get paper here
  • Baird, S.F. and Girard,C. 1852. Characteristics of some new reptiles in the Museum of the Smithsonian Institution. Proc. Acad. Nat. Sci. Philadelphia 6: 68-70 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Bezy, R. L., P. C. Rosen, T. R. Van Devender, and E. F. Enderson. 2017. Southern distributional limits of the Sonoran Desert herpetofauna along the mainland coast of northwestern Mexico. Mesoamerican Herpetology 4(1): 138–167 - get paper here
  • Bonetti, Mathilde 2002. 100 Sauri. Mondadori (Milano), 192 pp. - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Bruekers, Jaco and Peek, Ron 2004. Veldwaarnemingen aan de zuidelijke woestijn-Padhagedis, Phrynosoma platyrhinos calidiarum (COPE 1896) in Nevada en Zuid-Californië. Lacerta 62 (3): 106-112 [erratum in 62 (4): 174] - get paper here
  • Burt, Charles E. 1933. Some lizards from the Great Basin of the West and adjacent areas, with comments on the status of various forms. American Midland Naturalist 14: 228-250 - get paper here
  • Cope, E.D. 1896. On two new species of lizards from Southern California. American Naturalist 30: 833-836 - get paper here
  • Cope, E.D. 1900. The crocodilians, lizards and snakes of North America. Ann. Rep. U.S. Natl. Mus. 1898: 153-1270 - get paper here
  • Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92
  • Gentry, A.F. 1885. A review of the genus Phrynosoma. Proc. Acad. Nat. Sci. Philad. (ser. 3) 37: 138-148 - get paper here
  • Heimes, P. 2022. LIZARDS OF MEXICO - Part 1 Iguanian lizards. Edition Chimaira, Frankfurt Am Main, 448 pp.
  • Jones, L.L. & Lovich, R.E. 2009. Lizards of the American Southwest. A photographic field guide. Rio Nuevo Publishers, Tucson, AZ, 568 pp. [review in Reptilia 86: 84] - get paper here
  • Klauber, Laurence M. 1935. The status of the Sonoran horned toad, Phrynosoma goodei Stejneger. Copeia 1935 (4): 178-179 - get paper here
  • Lara-Resendiz, Rafael Alejandro, Tereza Jezkova, Philip C. Rosen and Fausto Roberto Mendez-de la Cruz. 2014. Thermoregulation during the summer season in the Goode's Horned Lizard Phrynosoma goodei (Iguania: Phrynosomatidae) in Sonoran Desert. Amphibia-Reptilia 35 (2): 161-172 - get paper here
  • Leaché, Adam D. and Jimmy A. McGuire 2006. Phylogenetic relationships of horned lizards (Phrynosoma ) based on nuclear and mitochondrial data: Evidence for a misleading mitochondrial gene tree. Molecular Phylogenetics and Evolution 39 (3): 628-644 - get paper here
  • Lemos-Espinal JA, Smith GR, Rorabaugh JC 2019. A conservation checklist of the amphibians and reptiles of Sonora, Mexico, with updated species lists. ZooKeys 829: 131-160 - get paper here
  • Meyers, J.J.; Herrel, A. & Nishikawa, K. 2006. Morphological correlates of ant eating in horned lizards (Phrynosoma). Biological Journal of the Linnean Society 89: 13–24 - get paper here
  • Mosauer, Walter 1936. The reptilian fauna of sand dune areas of Vizcaino Desert and of northwestern Lower California. Occasional Papers of the Museum of Zoology, University of Michigan (329): 1-21 - get paper here
  • Mulcahy, Daniel G.; Allen W. Spaulding, Joseph R. Mendelson III & Edmund D. Brodie, Jr. 2006. Phylogeography of the Flat-tailed Horned Lizard (Phrynosoma mcallii) and systematics of the P. mcallii–platyrhinos mtDNA complex. Molecular Ecology 15(0): 1–20 - get paper here
  • Pianka, E. R. 1991. Phrynosoma platyrhinos. Girard: Desert Horned Lizard. Catalogue of American Amphibians and Reptiles 517: 1-4 - get paper here
  • Pianka, E. R.;Parker, W. S. 1975. Ecology of horned lizards: A review with special reference to Phrynosoma platyrhinos. Copeia 1975 (1): 141-162 - get paper here
  • Powell, G. L. and A. P. Russell 2023. The horns of horned lizards (Phrynosomatidae: Phrynosoma): the long road to their conceptualization as unique lacertilian features with individual identity. Bibliotheca Herpetologica 17(10):86–107 - get paper here
  • Reeder,T.W. & Montanucci,R.R. 2001. Phylogenetic analysis of the horned lizards (Phrynomomatidae: Phrynosoma): evidence from mitochondrial DNA and morphology. Copeia 2001 (2): 309-323 - get paper here
  • Smith, H.M. & Taylor,E.H. 1950. An annotated checklist and key to the reptiles of Mexico exclusive of the snakes. Bull. US Natl. Mus. 199: 1-253 - get paper here
  • Smith, Hobart M. 1939. An annotated list of the Mexican amphibians and reptiles in the Carnegie Museum. Annals of the Carnegie Museum 27: 311-320
  • Stebbins,R.C. 1985. A Field Guide to Western Reptiles and Amphibians, 2nd ed. Houghton Mifflin, Boston
  • Stejneger, L.H. 1893. Annotated list of the reptiles and batrachians collected by the Death Valley Expedition in 1891, with descriptions of new species. North American Fauna, No. 7: 159-228 (+ 14 plates + 4 maps) - get paper here
  • Turner, Frederick B.;Wauer, Roland H. 1963. A survey of the herpetofauna of the Death Valley area. Great Basin Naturalist 23 (3-4): 119-128 - get paper here
  • Werning, H. 2011. Phrynosoma platyrhinos. Wüstenkrötenechse. Reptilia (Münster) 16 (90): 47-50 - get paper here
  • Werning, H. 2014. Krötenechsen – eine (sehr gute) Laune der Natur. Reptilia (Münster) 19 (107): 16-23
  • Werning, Heiko 2012. Die Reptilien und Amphibien des Südwestens. Draco 13 (50): 18-60 - get paper here
  • Woodbury, Angus Munn 1928. The reptiles of Zion National Park. Copeia 1928 (166): 14-21 - get paper here
  • Zorn, H. 2004. Die Wüste im eigenen Haus. Reptilia (Münster) 9 (48): 41-44 - get paper here
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