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Phymaturus delheyi AVILA, PEREZ, PEREZ & MORANDO, 2011

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Higher TaxaLiolaemidae, Iguania, Sauria, Squamata (lizards)
Common Names 
SynonymPhymaturus delheyi AVILA, PÉREZ, PEREZ & MORANDO 2011
Phymaturus delheyi — LOBO et al. 2021 
DistributionArgentina (NW Patagonia)

Type locality: rocky environments of the northern Tromen Vol- cano massif, along Butacó Creek, on Provincial Road 37 (36° 59’ S, 69° 59’ W, 1810 m, datum = WGS 84), Pehuenches Department, Neuquén Province, Argentina.  
TypesHolotype: MLP S 2609, adult male; L.J. Avila, D.R. Pérez, and C.H.F. Pérez, collectors. 
DiagnosisDiagnosis: Phymaturus delheyi is a robust and medium sized member of the clade of Phymaturus lizards referred to as the patagonicus group by Etheridge (1995) because it has flat imbricate superciliaries, non-rugose dorsal scales on tail, and subocular usually not fragmented. Phymaturus delheyi can be distinguished from other species of the patagonicus group (P. agilis, P. calcogaster, P. castillensis, P. ceii, P. desuetus, P. etheridgei, P. excelsus, P. felixi, P. indistinctus, P. manuelae, P. patagonicus, P. somuncurensis, P. spurcus, P. spectabilis, P. videlai, P. tenebrosus) by colour pattern features and geographical distribution. This new species is allopatric and differs from all other members of the clade in its unique dorsal pattern of medium size white spots covering 1–10 scales on a dark-brown background. The presence of sexual dichromatism differentiates Phymaturus delheyi from Phymaturus sitesi sp. nov. and P. nevadoi. Dorsal pattern of P. payuniae is composed by irregular white spots (between 4–40 scales each, sometimes fused) scattered along head and trunk, and becoming enlarged on tail to almost reticulated on limbs, a pattern never observed in P. delheyi. Ventral color- ation in P. payuniae is clearer than in P. delheyi. Phymaturus delheyi has a pink tinge on the chest and belly, becoming an orange coloration on the lower abdomen, and cloacal and femoral regions not observed in P. payuniae. Scales around midbody are higher in P. payuniae than in P. delheyi and show little overlap (221–234 vs 198–227). Dorsal pattern of P. zapalensis is composed by irregular white spots (between 5–14 scales each, sometimes fused) scattered along head and trunk but not in limbs and tail, white spots become smaller in lateral areas, and then lateral dark areas turn into dark bands between shoulder and rump, a pattern never observed in P. delheyi. Ventral color- ation in P. zapalensis is greyish with a dark reticulated only in the gular region, pattern never observed in P. delheyi. Phymaturus nevadoi has a dorsal pattern with spots usually occupying fewer scales (4–9 scales vs 1–10), greater density, and grow to be larger on flanks (not in P. delheyi). Phymaturus delheyi shows a higher ventral scale count (174–202 vs 162–179 in P. nevadoi). Phymaturus delheyi can be distinguished from Phymaturus sitesi new sp. by the dorsal pattern described above. Scales around midbody are fewer in P. delheyi than in P. sitesi (198–227 vs 210–229), and higher in ventral scales with some overlap (174–202 vs 160–190). 
CommentAbundance: only known from its original description (Meiri et al. 2017). 
Etymology“The species is dedicated to honor our friend Kaspar Delhey from Max Planck Institute for Ornithology, Germany, an Argentinean ornithologist and evolutionary biologist who helped to collected this and other species still to be described from northern Patagonia.” 
  • AVILA, LUCIANO JAVIER; CRISTIAN HERNAN FULVIO PEREZ, DANIEL ROBERTO PEREZ & MARIANA MORANDO 2011. Two new mountain lizard species of the Phymaturus genus (Squamata: Iguania) from northwestern Patagonia, Argentina. Zootaxa 2924: 1–21 - get paper here
  • Avila, Luciano Javier; Cristian Hernan Fulvio Perez, IGNACIO MINOLI & Mariana Morando 2014. A new lizard of the Phymaturus genus (Squamata: Liolaemidae) from Sierra Grande, northeastern Patagonia, Argentina. Zootaxa 3793 (1): 099–118 - get paper here
  • AVILA, LUCIANO JAVIER; LORENA ELIZABETH MARTINEZ & MARIANA MORANDO 2013. Checklist of lizards and amphisbaenians of Argentina: an update. Zootaxa 3616 (3): 201–238 - get paper here
  • GONZÁLEZ-MARÍN, ANDREA; CRISTIAN hERNÁN FULVIO PÉREZ, IGNACIO MINOLI, Mariana Morando, Luciano Javier Avila 2016. A new lizard species of the Phymaturus patagonicus group (Squamata: Liolaemini) from northern Patagonia, Neuquén, Argentina. Zootaxa 4121 (4): 412–430 - get paper here
  • Lobo, F., Barrasso, D. A., Hibbard, T., Quipildor, M., Slodki, D., Valdecantos, S., & Basso, N. G. 2021. Morphological and genetic divergence within the Phymaturus payuniae clade (Iguania: Liolaemidae), with the description of two new species. South American Journal of Herpetology 20 (1): 42-66 - get paper here
  • Lobo, Fernando; Santiago Javier Nenda 2015. Discovery of two new species of Phymaturus (Iguania: Liolaemidae) from Patagonia, Argentina, and occurrence of melanism in the patagonicus group. Cuad. herpetol. 29 (1): 5-25
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
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