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Pituophis melanoleucus (DAUDIN, 1803)

IUCN Red List - Pituophis melanoleucus - Least Concern, LC
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Higher TaxaColubridae, Colubrinae, Lampropeltini, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
SubspeciesPituophis melanoleucus lodingi BLANCHARD 1924
Pituophis melanoleucus melanoleucus (DAUDIN 1803)
Pituophis melanoleucus mugitus BARBOUR 1921 
Common NamesE: Eastern Pine Snake
lodingi: Black Pine Snake
melanoleucus: Northern Pine Snake
mugitus: Florida Pine Snake; G: Florida-Kiefernnatter
G: Bullennatter 
SynonymColuber melanoleucus DAUDIN 1803: 188
Pituophis melanoleucus — HOLBROOK 1842: 7
Churchillia bellona BAIRD & GIRARD 1852: 70
Pituophis bellona — BAIRD & GIRARD 1852: 351
Pituophis melanoleucus — BAIRD & GIRARD 1853: 65
Rhinechis melanoleucus — DUMÉRIL 1853: 453
Pituophis melanoleucus — DUMÉRIL & BIBRON 1854: 233
Pityophis melanoleucus — GARMAN 1884: 51
Pityophis catenifer var. bellona — GARMAN 1884: 53
Pityophis melanoleucus — SCLATER 1891: 30
Coluber melanoleucus — BOULENGER 1894: 68
Pityophis melanoleucus — COPE 1900: 867
Pityophis sayi bellona — COPE 1900: 872
Pituophis melanoleucus — STULL 1940
Pituophis melanoleucus — SMITH & SMITH 1976
Pituophis melanoleucus — STEBBINS 1985: 189
Pituophis melanoleucus — CONANT & COLLINS 1991: 202
Pituophis melanoleuca — BLUME 1999
Pituophis melanoleucus — ERNST & ERNST 2003: 280
Pituophis melanoleucus — BURBRINK 2007
Pituophis melanoleucus — WALLACH et al. 2014: 559

Pituophis melanoleucus lodingi BLANCHARD 1924
Pituophis lödingi BLANCHARD 1924
Pituophis melanoleucus loedingi — STULL 1940: 79
Pituophis melanoleucus loedingi — KLUGE 1984
Pituophis melanoleucus lodingi — CROTHER 2000
Pituophis melanoleucus lodingi — TENNANT & BARTLETT 2000: 364
Pituophis melanoleucus lodingi — BAXLEY et al. 2011
Pituophis melanoleucus lodingi — SCHÄBERLE 2012

Pituophis melanoleucus melanoleucus (DAUDIN 1803)
Coluber melanoleucus DAUDIN 1803: 188
Pituophis melanoleucus melanoleucus — STULL 1940
Pituophis melanoleucus melanoleucus — TENNANT & BARTLETT 2000: 360
Pituophis melanoleucus melanoleucus — CROTHER 2000
Pituophis melanoleucus melanoleucus — GUYER et al. 2018

Pituophis melanoleucus mugitus BARBOUR 1921
Pituophis melanoleucus mugitus — STULL 1940
Pituophis melanoleucus mugitus — CROTHER 2000
Pituophis melanoleucus mugitus — TENNANT & BARTLETT 2000: 363
Pituophis melanoleucus mugitus — HIRSCHKORN & SKUBOWIUS 2011
Pituophis melanoleucus mugitus — GUYER et al. 2018 
DistributionUSA (Louisiana, Kentucky, Tennessee, Virginia, North Carolina, South Carolina, Georgia, Alabama, Mississippi, Florida, New Jersey)

melanoleucus: USA (Virginia, S North Carolina west through South Carolina to northern Georgia, Tennessee, Kentucky, south into Alabama. Disjunct populations in S New Jersey, WC Virginia, adjacent West Virginia, C Kentucky, SW Tennessee). Intergrades with mugitus in South Carolina, Georgia, and Alabama. Type locality: "la Caroline méridionale et de la Floríde." Restricted to Carolina by Barbour, 1921b: 117. Further restricted to Charleston, South Carolina, USA by Schmidt, 1953a: 201.

mugitus: USA (S South Carolina to Georgia and S Florida, extreme E and S Alabama); Type locality: Palm Beach, Florida, 26.705639° -80.036419° (WGS84), Error: 8000 m.

lodingi: USA (SW Alabama to E Louisiana, NW Florida). Type locality: Alabama, Mobile, between Irvington and Grand Bay.  
Reproductionoviparous 
TypesHolotype: nondesignated, present location unknown.
Holotype: UMMZ 58800 [lodingi]
Holotype: MCZ R-15525 [mugitus]
Holotype: USNM 1519 [Churchillia bellona] 
DiagnosisAdditional details, e.g. a detailed description or comparisons (7507 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSubspecies: Pituophis melanoleucus annectens, P. m. sayi, and other subspecies have been transferred to P. catenifer. Not listed in LINER 1994. Nikolakis et al. 2021 find that the current subspecies do not appear to represent independent lineages, but, rather, show a significant amount of admixture across the entire P. melanoleucus range. Khakurel et al. 2023 do not find the traditionally recognized subspecies to be monophyletic and speculate that there may be as many as 8 different clades.

Distribution: map for subspecies in Rodríguez-Robles & De Jesús-Escobar (2000). These authors also found that two segments of the melanoleucus complex, the lodingi–melanoleucus–mugitus eastern pinesnake clade and the affinis–annectens–bimaris–catenifer–deserticola–sayi–ruthveni–vertebralis clade from central and western United States and northern Mexico, represent divergent, allopatric lineages with no known intergradation zone. Rodríguez-Robles & De Jesús-Escobar recognize each of these two groupings as a different species (see P. catenifer). Not in Mexico (Heimes 2016). See map in Khakurel et al. 2023: Fig. 1.

Type species: Coluber melanoleucus DAUDIN 1803: 188 is the type species of the genus Pituophis HOLBROOK 1842.

Phylogenetics: see Dahn et al. 2018 for a phylogeny of Lampropeltini. 
EtymologyThe genus name was derived from the Greek words pitys, meaning "pine" and ophis, meaning "snake." 
References
  • Baird, S. F. and C. Girard. 1853. Catalogue of North American Reptiles in the Museum of the Smithsonian Institution. Part 1.-Serpents. Smithsonian Inst., Washington, xvi + 172 pp. - get paper here
  • Baird, S.F. and Girard,C. 1852. Characteristics of some new reptiles in the Museum of the Smithsonian Institution. Proc. Acad. Nat. Sci. Philadelphia 6: 68-70 - get paper here
  • Baird,S.F. and Girard,C. 1852. Reptiles. in: Stansbury’s Exploration and survey of the valley of the Great Salt Lake of Utah, 1852, including a Reconnoissance of a new Route through the Rocky Mountains. Philadelphia, Lippincott & Co., pp. 336-365 - get paper here
  • Barbour, R. 1921. The Florida Pine snake. Proc. New England Zool. Club 7: 117-118 - get paper here
  • Barbour, Thomas 1940. Pine snakes, black and brown. Copeia 1940 (3): 205 - get paper here
  • Baxley, D.L. & C.P. Qualls 2009. Black Pine Snake (Pituophis melanoleucus lodingi): Spatial Ecology and Associations between Habitat Use and Prey Dynamics Journal of Herpetology 43 (2): 284-293. - get paper here
  • Baxley, Danna; Gregory J Lipps, and Carl P Qualls 2011. Multiscale Habitat Selection by Black Pine Snakes (Pituophis melanoleucus lodingi) in Southern Mississippi. Herpetologica 67 (2): 154-166. - get paper here
  • Beane, J. C. 2012. Pituophis melanoleucus melanoleucus (northern pinesnake) male-male combat. Herpetological Review 43: 349-350 - get paper here
  • Bechtel, Elizabeth R.;Whitecar, Terrence 1983. Genetics of striping in the gopher snake, Pituophis melanoleucus. Journal of Herpetology 17 (4): 362-370 - get paper here
  • Bechtel, H.B., J.W. Nelson & E. Bechtel 1980. Histochemical Demonstration of Two Types of Albinism in San Diego Gopher Snakes (Pituophis melanoleucus annectens) by Use of Dopa Reaction Copeia 1980 (4): 932-935. - get paper here
  • Blackburn, D.G. 1994. Standardized criteria for the recognition of embryonic nutritional patterns in squamate reptiles. Copeia 1994 (4): 925-935 - get paper here
  • Blanchard, F.N. 1924. A name for the Black Pituophis from Alabama. Papers Michigan Acad.Sci, Arts Letters 4: 531-532
  • Blanchard, Frank N. 1920. A Black Pituophis. Copeia 1920 (81): 30-33 - get paper here
  • Blume, F. 1999. Interessantes Drohverhalten einer Kiefernnatter, Pituophis melanoleuca (Daudin, 1803) im Sandhill Game Land Nationalpark, North Carolina (U. S. A.). Sauria 21 (1): 45-46 - get paper here
  • Bogosian, Victor; Eric C. Hellgren, and Raymond W. Moody 2012. Assemblages of Amphibians, Reptiles, and Mammals on an Urban Military Base in Oklahoma. Southwestern Naturalist 57 (3): 277-284. - get paper here
  • Boulenger, George A. 1894. Catalogue of the snakes in the British Museum (Natural History). Volume II., Containing the Conclusion of the Colubridæ Aglyphæ. British Mus. (Nat. Hist.), London, xi, 382 pp. - get paper here
  • Brimley, C.S. 1918. Pituophis melanoleucus in North Carolina Copeia 63: 92. - get paper here
  • Bryant, C.J. 1982. Literature: Breeding the Gopher snake, Pituophis melanoleucus. Litteratura Serpentium 3 (5): 260 - get paper here
  • Burger J, Zappalorti RT, Gochfeld M 2018. Hatchling survival to breeding age in Northern Pine Snakes (Pituophis melanoleucus) in the New Jersey Pine Barrens: Human effects on recruitment from 1986 to 2017. PLoS ONE 13(5): e0195676 - get paper here
  • Burger J. 2019. Vulnerability of Northern Pine Snakes (Pituophis melanoleucus Daudin, 1803) during fall den ingress in New Jersey, USA. Amphibian & Reptile Conservation 13(2) [General Section]: 102–114 (e191) - get paper here
  • Burger, J. 1990. Response of Hatchling Pine Snakes (Pituophis melanoleucus) to Chemical Cues of Sympatric Snakes Copeia 1990 (4): 1160-1163. - get paper here
  • Burger, J. & R.T. Zappalorti 1986. Nest Site Selection by Pine Snakes, Pituophis melanoleucus, in the New Jersey Pine Barrens Copeia 1986 (1): 116-121. - get paper here
  • Burger, J. & R.T. Zappalorti 1988. Habitat Use in Free-Ranging Pine Snakes, Pituophis melanoleucus, in New Jersey Pine Barrens Herpetologica 44 (1): 48-55. - get paper here
  • Burger, J. & R.T. Zappalorti 1989. Habitat use by Pine Snakes (Pituophis melanoleucus) in the New Jersey Pine Barrens: Individual and Sexual Variation Journal of Herpetology 23 (1): 68-73. - get paper here
  • Burger, J., R.T. Zappalorti, J. Dowdell, T. Georgiadis, J. Hill & M. Gochfeld 1992. Subterranean Predation on Pine Snakes (Pituophis melanoleucus) Journal of Herpetology 26 (3): 259-263. - get paper here
  • Burger, J., R.T. Zappalorti, M. Gochfeld, W.I. Boarman, M. Caffrey, V. Doig, S.D. Garber, B. Lauro, M. Mikovsky, C. Safina & J. Saliva 1988. Hibernacula and Summer Den Sites of Pine Snakes (Pituophis melanoleucus) in the New Jersey Pine Barrens Journal of Herpetology 22 (4): 425-433. - get paper here
  • Burger, Joanna and Robert T. Zappalorti 2011. THE NORTHERN PINE SNAKE (PITUOPHIS MELANOLEUCUS): ITS LIFE HISTORY, BEHAVIOR AND CONSERVATION. Nova Science Publishers, NY, 92 pp.
  • Burger, Joanna and Robert Zappalorti 2015. Hibernation Site Philopatry in Northern Pine Snakes (Pituophis melanoleucus) in New Jersey. Journal of Herpetology Jun 2015, Vol. 49, No. 2: 245-251. - get paper here
  • Burger, Joanna, Robert T. Zappalorti and Michael Gochfeld. 2000. Defensive behaviors of Pine Snakes (Pituophis melanoleucus ) and Black Racers (Coluber constrictor) to disturbance during hibernation. Herpetological Natural History 7(1):59-66
  • Burger, Joanna; Robert T. Zappalorti, Michael Gochfeld, Emile DeVito, David Schneider, Matt McCort, and Christian Jeitner 2012. Long-Term Use of Hibernacula by Northern Pinesnakes (Pituophis melanoleucus). Journal of Herpetology 46 (4): 596-601. - get paper here
  • Burger, Joanna; Robert Zappalorti, Michael Gochfeld, Emile Devito & Christian Jeitner 2021. Trade-Offs and Vulnerability of Northern Pine Snakes (Pituophis m. melanoleucus): A Comparison of Nest-Site Selection in the Early 1980s and 2020. Herpetologica Sep 2021 Vol. 77, No. 3: 249-258 - get paper here
  • BURGER, JOANNA; ROBERT. T. ZAPPALORTI, MICHAEL GOCHFELD & EMILE DEVITO. 2021. The importance of paleodunes as nesting habitat for Northern Pine Snakes (Pituophis melanoleucus melanoleucus): risk from off-road vehicles in the New Jersey Pine Barrens. Urban Ecosystems (2021): 1–12. - get paper here
  • Camper, Jeffrey D. 2019. The Reptiles of South Carolina. University of South Carolina Press, 288 pp. [review in Copeia 107 (3): 590, 2019] - get paper here
  • Chermock, R.L. 1955. A Record of the Northern Pine-Snake, Pituophis melanoleucus melanoleucus, from Alabama Copeia 1955 (2): 141. - get paper here
  • Chiszar, D., V. Lipetz, K. Scudder & E. Pasanello 1980. Rate of Tongue Flicking by Bull Snakes and Pine Snakes (Pituophis melanoleucus) during Exposure to Food and Non-Food Odors Herpetologica 36 (3): 225-231. - get paper here
  • Cliburn, J.W. 1962. Further Notes on the Behavior of a Captive Black Pine Snake (Pituophis melanoleucus lodingi Blanchard) Herpetologica 18 (1): 34-37. - get paper here
  • Cliburn, J.W. 1975. The Hemipenis of Pituophis melanoleucus Journal of Herpetology 9 (2): 254-255. - get paper here
  • Cliburn, J.W. 1976. Observations of Ecdysis in the Black Pine Snake, Pituophis melanoleucus lodingi (Reptilia, Serpentes, Colubridae) Journal of Herpetology 10 (4): 299-301. - get paper here
  • Conant,R. & Collins,J.T. 1991. A Field Guide to Reptiles and Amphibians of Eastern/Central North America, 3rd ed. Houghton Mifflin (Boston/New York), xx + 450 p.
  • Cope, E.D. 1900. The crocodilians, lizards and snakes of North America. Ann. Rep. U.S. Natl. Mus. 1898: 153-1270 - get paper here
  • Cresswell, G.F. 1971. Albinism in the Northern Pine Snake, Pituophis melanoleucus melanoleucus Bull. Maryland Herpetol. Soc., 7(4): 91-93. - get paper here
  • Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92
  • Dahn, H. A., Strickland, J. L., Osorio, A., Colston, T. J., & Parkinson, C. L. 2018. Hidden diversity within the depauperate genera of the snake tribe Lampropeltini (Serpentes, Colubridae). Molecular Phylogenetics and Evolution - get paper here
  • Daudin, F. M. 1803. Histoire Naturelle Generale et Particuliere des Reptiles. Vol. 6. F. Dufart, Paris, 447 pp. - get paper here
  • De Queiroz, A. 1984. Effects of Prey Type on the Prey-handling Behavior of the Bullsnake, Pituophis melanoleucus Journal of Herpetology 18 (3): 333-336. - get paper here
  • Degenhardt, William G.; C. W. Painter, and A. H. Price 1996. Amphibians and reptiles of New Mexico. Univ. New Mexico Press, 431 pp.
  • Dekay 1842. Fauna, New York, Rept.
  • Dodd, C K, Jr.; 2019. Pituophis melanoleucus mugitus (Florida Pine Snake). Longevity and tumor development. Herpetological Review 50 (3): 519 - get paper here
  • Echternacht, A.C. 1964. Albino specimens from Arizona Herpetologica 20 (3): 211-212. - get paper here
  • Eerden, L.J.A. van der 1989. Successful breeding of the Bullsnake (Pituophis melanoleucus sayi). Litteratura Serpentium 9 (3): 129-132 - get paper here
  • Elmberg, J. 1980. Herpetological observations 77-78 [in Vernon county, Wisconsin]. Fältbiologerna Umea.
  • Enge, Kevin M. 2009. Venomous and non-venomous snakes of Florida. Publication of the Florida Fish & Wildlife Conservation Commission. 16 pp.
  • Enge, Kevin M., Jonathan D. Mays and Cody D. Godwin. 2015. Geographic Distribution: Pituophis melanoleucus mugitus (Florida pinesnake). Herpetological Review 46 (2): 221 - get paper here
  • Ernst, C.H. & Ernst, E.M. 2003. Snakes of the United States and Canada. Smithsonian Inst. Press, Washington, 668 pp.
  • Ernst, C.H., & Barbour, R.W. 1989. Snakes of eastern North America. George Mason Univ. Press, Fairfax, VA 282 pp.
  • Fantuzzi, J. A., K. Jamieson, K. Pollack and B. Zarate 2016. Pituophis melanoleucus melanoleucus (North American Pinesnake) reproduction / ritualistic combat / courtship behavior. Herpetological Review 47(3): 481-482. - get paper here
  • Gates, G.O. 1955. A Record Length for the Arizona Bull Snake Pituophis melanoleucus affinis from Arizona Herpetologica 11 (3): 235. - get paper here
  • Gerald, G.W., M.A. Bailey & J.N. Holmes 2006. Movements and Activity Range Sizes of Northern Pinesnakes (Pituophis melanoleucus melanoleucus) in Middle Tennessee Journal of Herpetology 40 (4): 503-510. - get paper here
  • GODLEY, J. S., J. L. HOFKES, AND C. M. REILLY 2020. Minimum Density Estimates of Terrestrial Snakes in the Munson Sandhills Adjoining Apalachicola National Forest, Florida. Herpetological Review 51: 221 - 226.
  • González-Romero, A., & Alvarez-Cárdenas, S. 1989. Herpetofauna de la Region del Pinacate, Sonora, Mexico: Un Inventario. The Southwestern Naturalist, 34(4), 519–526 - get paper here
  • Graham, Sean P. 2011. Geographic Distribution: Pituophis melanoleucus (pinesnake). Herpetological Review 42 (3): 395 - get paper here
  • Green, N.B., & Pauley, T.K. 1987. Amphibians and reptiles in West Virginia. Univ. of Pittsburgh Press, Pittsburgh, 241 pp.
  • Grogan, W. L. and C. M. Heckscher. 2001. Are northern pine snakes, Pituophis m. melanoleucus, indigenous to Delaware? The Maryland Naturalist 44:20-36.
  • Grogan, W.L., Jr. 1973. A Northern Pine snake, Pituophis m. melanoleucus, from Maryland Bull. Maryland Herpetol. Soc., 9(2): 27-30. - get paper here
  • Gutzke, W.H.N., G.L. Paukstis & L.L. McDaniel 1985. Skewed Sex Ratios for Adult and Hatchling Bullsnakes, Pituophis melanoleucus, in Nebraska Copeia 1985 (3): 649-652. - get paper here
  • Guyer, Craig; Mark A. Bailey, and Robert H. Mount 2018. Lizards and snakes of Alabama. University of Alabama Press, 397 pp. - get paper here
  • Hirschkorn, D. 1986. Breeding results: Pituophis melanoleucus affinis. Litteratura Serpentium 6 (4): 158 - get paper here
  • Hirschkorn, D. 1986. Breeding results: Pituophis melanoleucus catenifer. Litteratura Serpentium 6 (4): 157-158 - get paper here
  • Hirschkorn, D. 1986. Breeding results: Pituophis melanoleucus annectans. Litteratura Serpentium 6 (4): 158-159 - get paper here
  • Hirschkorn, D. 1987. Breeding results: Pituophis melanoleucus catenifer. Litteratura Serpentium 7 (1): 47 - get paper here
  • Hirschkorn, D. & Skubowius, B. 2011. Bullennattern, Gophernattern, Kiefernnattern - Die Gattung Pituophis. Natur und Tier-Verlag (NTV), Münster, 176 pp. [review in Reptilia 16 (91): 85] - get paper here
  • Hirschkorn, Dieter 1987. Erfahrungen bei der Pflege und Zucht der selten gehaltenen Bullennatter Pituophis melanoleucus catenifer. Litteratura Serpentium 7 (4): 198-199 - get paper here
  • Holland, A; Enge, K M & Talley, B L; 2019. Pituophis melanoleucus mugitus (Florida Pinesnake) Reproduction/nest site. Herpetological Review 50 (2): 398-399 - get paper here
  • Hudson, Bryan D., Zach I. Felix, Justin Oguni, Brad Wilson, Kira McEntire, Theresa Stratmann, Daniel D. Duff and Zack Seymour. 2015. New geographic distributional records of amphbians and reptiles in Georgia, USA. Herpetological Review 46 (4): 595-596 - get paper here
  • Irwin, Kelly J. 2004. Arkansas Snake Guide. Arkansas Game & Fish Commission, Little Rock, 50 pp.
  • Jensen, John B.; Carlos D. Camp, Whit Gibbons, & Matt J. Elliott 2008. Amphibians and reptiles of Georgia. University of Georgia Press, 575 pp.
  • Klauber, L. M. 1947. Classification and ranges of the gopher snakes of the genus Pituophis in the United States. Bulletins of the Zoological Society of San Diego (22): 7-81 - get paper here
  • Klauber, Laurence M. 1946. The gopher snakes of Baja California, with descriptions of new subspecies of Pituophis catenifer. Transactions of the San Diego Society of Natural History 11 (1): 1-40 + 2 plates - get paper here
  • Knight, James L. 1986. Variation in snout morphology in the North American snake Pituophis melanoleucus (Serpentes: Colubridae). Journal of Herpetology 20 (1): 77-79 - get paper here
  • Landreth, H.F. 1972. Physiological Responses of Elaphe obsoleta and Pituophis melanoleucus to Lowered Ambient Temperatures Herpetologica 28 (4): 376-380. - get paper here
  • Lee, D.S. 1967. Eggs and Hatchlings of the Florida Pine Snake, Pituophis melanoleucus mugitus Herpetologica 23 (3): 241-242. - get paper here
  • Lee, J. R., et al. 2017. Geographic Distribution: Pituophis melanoleucus lodingi (Black Pine Snake). Herpetological Review 48 (2): 392 - get paper here
  • Leonard, William P.;Phenix, Erik 1994. Pituophis melanoleucus (gopher snake). USA: Washington. Herpetological Review 25 (4): 168 - get paper here
  • Liner, E.A. 1994. Scientific and common names for the Amphibians and Reptiles of Mexico in English and Spanish. Herpetological Circular 23: 1-113
  • Linzey, D.W., & Clifford, M.J. 1981. Snakes of Virgina. Univ. Press of Virginia, Charlottesville 159 pp.
  • Lönnberg, Einar 1894. Notes on reptiles and batrachians collected in Florida in 1892 and 1893. Proc. US Natl. Mus. 17 (1003): 317-339 - get paper here
  • Martin, W.F. & R.B. Huey 1971. The function of the epiglottis in sound production (hissing) of Pituophis melanoleucus Copeia 1971 (4): 752-754. - get paper here
  • Maxwell, John, Jr. and A. Floyd Scott. 2014. Geographic Distribution: Pituophis melanoleucus melanoleucus (northern pinesnake). Herpetological Review 45 (3): 466 - get paper here
  • McKelvy, Alexander D. and Alex Figureoa. 2014. Geographic Distribution: Pituophis melanoleucus (pinesnake). Herpetological Review 45 (1): 96 - get paper here
  • McKeown, Sean;Morgan, Mary 1996. Two size records for California snakes. Bull. Chicago Herp. Soc. 31 (12): 217-218 - get paper here
  • Mehrtens, J.M. 1952. Notes on the Eggs and Young of Pituophis melanoleucus sayi Herpetologica 8 (3): 69-70. - get paper here
  • Miller, Gabriel J.; Lora L. Smith, Steve A. Johnson, and Richard Franz 2012. Home Range Size and Habitat Selection in the Florida Pine Snake (Pituophis melanoleucus mugitus). Copeia Dec 2012, Vol. 2012, No. 4: 706-713. - get paper here
  • Mitchell, J. C. & Reay, K.K. 1999. Atlas of amphibians and reptiles in Virginia. Specialty Publication 1, VA Dept of Game and Fisheries, 122 pp. - get paper here
  • Mitchell, J.C. 1994. The reptiles of Virginia. Virginia Department of Game and Inland Fisheries, ca. 350 pp.
  • Neill, W.T. 1947. Range of Pituophis m. melanoleucus Restricted Herpetologica 4 (1): 29-30. - get paper here
  • New, J.G. 1953. The Young of the Snake Pituophis melanoleucus melanoleucus Copeia 1953 (3): 183-184. - get paper here
  • Nikolakis, Zachary L.; Richard W. Orton, Brian I. Crother 2021. Fine-scale population structure within an Eastern Nearctic snake complex (Pituophis melanoleucus). Zoologica Scripta 51(2): 133–146 [online 2021, print 2022] - get paper here
  • Orton, R W; Hayman, B & Wallrichs, M; 2018. Pituophis melanoleucus mugitus (Florida Pinesnake) Refugia fidelity. Herpetological Review 49 (4): 762 - get paper here
  • Palmer, W.M. & Braswell, A.L. 1995. Reptiles of North Carolina. Univ. North Carolina Press
  • Perez-Higareda 1981. The occurrence of Pituophis melanoleucus sayi in Veracruz (Serpentes: Colubridae). Bull. Maryland Herp. Soc. 17 (2): 64-67 - get paper here
  • Pyron, R.Alexander; Frank T. Burbrink 2009. Neogene diversification and taxonomic stability in the snake tribe Lampropeltini (Serpentes: Colubridae). Molecular Phylogenetics and Evolution 52 (2): 524-529 - get paper here
  • Repp, R. 2002. Sonoran Gopher Snake (Pituophis melanoleucus affinis). Herpetofauna of the 100 Mile Circle. Sonoran Herpetologist 15 (6):68. - get paper here
  • Reynolds, R.P. & Scott Jr., N.J. 1982. Use of a Mammalian Resource by a Chihuahuan Snake Community. In: Scott, N.J. (ed.) Herpetological communities [an SSAR symposium]. US Fish and Wildlife Service, Wildl. Res. Rep. 13: 99-118
  • Rodríguez-Robles,J.A. & De Jesús-Escobar,J.M. 2000. Molecular Systematics of New World Gopher, Bull, and Pinesnakes (Pituophis: Colubridae), a transcontinental Species Complex. Molecular Phylogenetics and Evolution 14 (1): 35-50 - get paper here
  • Rushton, E. & Rushton, C. 2012. Geographic distribution: Pituophis melanoleucus mugitus (Florida pinesnake). Herpetological Review 43: 449-450 - get paper here
  • Savage, Jay M. 1948. The pine snake in Kentucky. Copeia 1948 (4): 302 - get paper here
  • Schäberle, W. 2012. Die Historie der Indigonatter in der Terraristik. Ophidia 6 (2): 25-29 - get paper here
  • Sclater, W. L. 1891. List of snakes in the Indian Museum. Trustees of the Indian Museum, Calcutta, 79 pp. - get paper here
  • Scott, A. Floyd 2015. Erratum for Pituophis m. melanoleucus in HR45(4):466. Herpetological Review 46 (1): 64 - get paper here
  • Shreve, B. 1945. Pituophis melanoleucus mugitus in Alabama Copeia 1945 (4): 234. - get paper here
  • Skubowius, B. 2009. Die Florida-Kiefernnatter, Pituophis melanoleucus mugitus BARBOUR, 1921 - bisher noch ein seltener Terrarienpflegling. Draco 10 (37): 64-71 - get paper here
  • Skubowius, B. 2009. Die Nördliche Kiefernnatter (Pituophis melanoleucus melanoleucus) in Natur und Terrarium. Teil 2 Terraria (20) 4 (6): 50-58.
  • Skubowius, B. 2009. Die Nördliche Kiefernnatter (Pituophis melanoleucus melanoleucus) in Natur und Terrarium. Teil 1 Terraria (19) 4 (5): 54-60.
  • Skubowius, B. 2010. New Jersey – Mit „Fieldherpern“ und „Pineys“ auf der Suche nach der Nördlichen Kiefernnatter (Pituophis melanoleucus melanoleucus). Reptilia (Münster) 15 (84): 52-60 - get paper here
  • Skubowius, Bernd 2004. Keeping and breeding Pine Snakes Pituophis melanoleucus. Litteratura Serpentium 24 (2): 67-77 - get paper here
  • Skubowius, Bernd 2008. Auf den Spuren von Carl Kauffeld durch New Jersey – nicht nur der Kiefernattern, Pituophis m. melanoleucus (Daudin, 1803) wegen. Ophidia 2 (1): 18-29 - get paper here
  • Skubowius, Bernd 2014. Bullige Schönheiten. Farb- und Zeichnungsformen bei Pituophis. Draco 16 (57): 40-47 - get paper here
  • Smith, Hobart M.;Thompson, Dorian 1993. Four reptiles newly recorded from Ouray County, Colorado. Bull. Chicago Herp. Soc. 28 (4): 78-79 - get paper here
  • Smith, Ronald M.; James R. Spotila, and Walter F. Bien 2015. Spatial Ecology of Northern Pinesnakes (Pituophis m. melanoleucus) in Disturbed and Undisturbed Habitats in the New Jersey Pine Barrens. Herpetologica Mar 2015, Vol. 71, No. 1: 19-25. - get paper here
  • Smith,H.M. & Smith,R.B. 1976. Synopsis of the Herpetofauna of Mexico. Vol. III. Source analysis and index for Mexican reptiles. John Johnson, North Bennington, Vermont.
  • Stebbins,R.C. 1985. A Field Guide to Western Reptiles and Amphibians, 2nd ed. Houghton Mifflin, Boston
  • STEVENSON, D. J 2020. Pituophis melanoleucus mugitus (Florida Pinesnake). Mortality. Herpetological Review 51:
  • Stevenson, D.J. 2017. Pituophis melanoleucus mugitus (Florida Pinesnake) Reproduction / Nest Site. Herpetological Review 48 (3): 680. - get paper here
  • STEVENSON, DIRK J. et al. 2021. New County Records for Amphibians and Reptiles in Georgia, USA. Herpetological Review 52 (2): 350-359
  • Stull, Olive Griffith 1940. Variations and relationships in the snakes of the genus Pituophis. Bull. US Natl. Mus. (175): 1-225 - get paper here
  • Stull, Olive Griffth 1932. An annotated list of the forms of the genus Pituophis. Occasional Papers of the Museum of Zoology, University of Michigan (250): 1-5 - get paper here
  • Sweet, S S; Parker, W S 1990. Pituophis melanoleucus (Daudin). Pine, bull, and gopher snakes. Catalogue of American Amphibians and Reptiles ( 474: 1-8 - get paper here
  • Sweet, S.S. 1985. Geographic Variation, Convergent Crypsis and Mimicry in Gopher Snakes (Pituophis melanoleucus) and Western Rattlesnakes (Crotalus viridis) Journal of Herpetology 19 (1): 55-67. - get paper here
  • Tanner, Vasco M.;Hayward, C. Lynn 1934. A biological study of the La Sal Mountains, Utah report No. 1 (Ecology). Proceedings of the Utah Academy of Sciences, Arts, and Letters 11: 209-235
  • Tennant, A. 2003. Snakes of North America - Eastern and Central Regions, revised edition. Lone Star Books, 605 pp.
  • Tennant, A. & Bartlett, R.D. 2000. Snakes of North America - Eastern and Central Regions. Gulf Publishing, Houston, TX, 588 pp.
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Werler, John E. & James R. Dixon 2000. Texas Snakes. University of Texas Press, 544 pages
  • Werning, H. 2009. Florida - ein kleiner herpetologischer Reiseführer. Draco 10 (37): 4-23 - get paper here
  • Wood, J.T. 1947. Pituophis m. melanoleucus in the Great Smoky Mountains National Park Area Herpetologica 3 (5): 152. - get paper here
  • Wood, Kristin N. 1998. Geographic distribution. Pituophis melanoleucus mugitus. Herpetological Review 29 (4): 249 - get paper here
  • Woodbury, Angus M.;Hansen, Richard M. 1950. A snake den in Tintic Mountains, Utah. Herpetologica 6: 66-70 - get paper here
  • Woodbury, Angus Munn 1928. The reptiles of Zion National Park. Copeia 1928 (166): 14-21 - get paper here
  • Wright, Johnathan 1996. Variations within populations of the bullsnake. Reptile & Amphibian Magazine 1996 (Sept./Oct.): 16-22
  • Zappalorti, R.T. 2012. Into the barrens. HerpNation 2.5 (8): 26-38 - get paper here
  • Zappalorti, Robert; Joanna Burger, and Frank Peterson 2015. Home Range Size and Distance Traveled from Hibernacula in Northern Pinesnakes in the New Jersey Pine Barrens. Herpetologica Mar 2015, Vol. 71, No. 1: 26-36. - get paper here
 
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