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Platyceps atayevi (TUNIYEV & SHAMMAKOV, 1993)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common Names 
SynonymColuber atayevi TUNIYEV & SHAMMAKOV 1993: 2
Platyceps atayevi — NAGY et al. 2004
Platyceps najadum atayevi — SCHÄTTI et al. 2005
Platyceps najadum atayevi — VENCHI & SINDACO 2006: 286
Platyceps atayevi — WALLACH et al. 2014: 564
Platyceps atayevi — ZAHER et al. 2019 
DistributionTurkmenistan (Kopet-Dag Mountains), probably Iran.

Type locality: “environs of Saivan Village, Saivan-Nokhur Plateau, western Kopet-Dag, Bakharden Region,Turkmenistan”  
Reproductionoviparous 
TypesHolotype: SNP (given as Collection of the Caucasian Reserve, Sochi, Russia), No. 420, adult male, 12 May 1991, collected by B. S. Tuniyev; paratypes: SNP, CAS 
DiagnosisDiagnosis: “Comparatively small snake (Fig. 2), smaller than C. n. najadum, C. n. dahli, and C. r. rubriceps in dimensions. It is comparable in size with the European Subspecies, C. rubriceps thracius. In contrast to C. najadum, whose tail constitutes 1/3 of its total length, C. atayevi has a comparatively short tail, approximately 1/4 of total length, also characteristic of C. rubriceps. Habitus and elements of coloration of C. atayevi are intermediate between C . najadum and C. rubriceps; its narrow, sharp, and flat head with the rostrum beveled downward is closer to the C. rubriceps head shape than to that of C. najadum, with its comparatively wide, rounded, and high head, where the upper and lower surfaces of head are parallel. Color pattern of C. atayevi resembles that of C. najadum, but brown colors prevail instead of olive-green ones. Lateral abdominal ridges are practically absent in C. atayevi, in contrast to the above mentioned species, and consequently the body is round in cross-section and not rectangular as in C. najadum and C. rubriceps. Genial scutes of C. atayevi contact one another along the mental groove, rarely a having a few isolated granules between the posterior pair, whereas between the widely separated posterior genials of C. najadum there are always 2-4 rows of well developed scales (Fig. 3). Posterior upper labials of C. atayevi are weakly distinguished from the throat scales, whereas all upper labials of C. najadum are strongly pronounced.” (TUNIYEV & SHAMMAKOV 1993) 
CommentSynonymy: Coluber atayevi has been synonymized with C. najadum by TOKAR et al. (1995) but considered as a valid species by more recent authors.

Similar species: See also P. najadum. 
EtymologyNamed after Turkmen herpetologist, Chary Ataevich Ataev, who studies reptiles of the mountains of Turkmenistan. 
References
  • Nagy, Z.T.; R. Lawson, U. Joger, M. Wink 2004. Molecular systematics of racers, whipsnakes and relatives (Reptilia: Colubridae) using mitochondrial and nuclear markers. Journal of Zoological Systematics and Evolutionary Research 42 (3): 223-233 - get paper here
  • Schatti, B., Stutz,A. & Chavret, C. 2005. Morphologie, Verbreitung und Systematik der Schlanknatter Platyceps najadum (Eichwald, 1831) (Reptilia: Squamata: Colubrinae). Revue Suisse de Zoologie 112 (3): 573-625 - get paper here
  • Tuniyev B S; Shammakov S M 1993. Coluber atayevi sp. nov. (Ophidia, Colubridae) from the Kopet-Dag Mountains of Turkmenistan. Asiatic Herpetological Research 5: 1-10 - get paper here
  • Venchi, Alberto and Roberto Sindaco 2006. Annotated checklist of the reptiles of the Mediterranean countries, with keys to species identification. Part 2 -Snakes (Reptilia, Serpentes). Annali del Museo Civico di Storia Naturale "G. Doria", Genova, XCVIII: 259-364
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Zaher H, Murphy RW, Arredondo JC, Graboski R, Machado-Filho PR, Mahlow K, et al. 2019. Large-scale molecular phylogeny, morphology, divergence-time estimation, and the fossil record of advanced caenophidian snakes (Squamata: Serpentes). PLoS ONE 14(5): e0216148 - get paper here
 
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