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Pristurus flavipunctatus RÜPPELL, 1835

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Higher TaxaSphaerodactylidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesRüppell’s Semaphore Gecko, Middle Eastern Rock Gecko 
SynonymPristurus flavipunctatus RÜPPELL 1835
Gymnodactylus flavipunctatus — DUMÉRIL & BIBRON 1836: 417
Pristurus flavipunctatus — BOULENGER 1885: 52
Pristurus percristatus BOULENGER 1896 (fide LOVERIDGE 1947)
Pristurus percristatus pseudoflavipunctatus SCORTECCI 1933 (fide LOV. 1947)
Pristurus flavipunctatus — LOVERIDGE 1947: 77
Pristurus flavipunctatus flavipunctatus — HAAS 1951
Pristurus gasperetti ARNOLD 1986 (fide SCHÄTTI & GASPERETTI 1994)
Pristurus gasperetti — KLUGE 1993
Pristurus flavipunctatus — KLUGE 1993
Pristurus flavipunctatus — RÖSLER 2000: 106
Pristurus gasperetti — RÖSLER 2000: 106
Pristurus flavipunctatus — LARGEN & SPAWLS 2010: 331 
DistributionE Ethiopia, Saudi Arabia (incl. Farasan Islands), Egypt, Djibouti, Somalia
Sudan (Neberet Valley),
Eritrea (Ghinda and Emberemi),
Jordan (Sandstone Hills)

Type locality: Massaua, Abessinien.

gasperetti (invalid): was described from W Saudi-Arabia (Wadi Kharrar near Makkah); Type locality: Wadi Kharrar, near Makkah, Saudi Arabia, 21° 17'30"N 40° 06'30"E, 460 m elevation. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesType: BMNH 1946.8.20.31-42 (and possibly additional specimens).
Holotype: BMNH 1986.206, male; Paratypes: BMNH [gasperetti]
 
CommentType: MSNTO R2512, Syntype: UMMZ (percristatus)

Type species: Pristurus flavipunctatus RÜPPELL 1835 is the type species of the genus Pristurus RÜPPELL 1835.

Subspecies: Pristurus gasperetti gallagheri ARNOLD 1986 has been elevated to full species.

Diagnosis (genus): dorsal process of premaxilla relatively long, nasal bones short, extent of prefrontal bone in anterior wall of orbit reduced medially*; supratemporal process of parietal bone often expanded and fully attached to exoccipital*, squamosal often short and curved*; palatine contacting vomer narrowly*, superficial posterior margin of dentary bone not obviously tridentate, the lower spur extending far backwards*. Twenty-five or fewer presacral vertebrae; toes with primitive number of phalanges (manus – 2.3.4.5.3; pes – 2.3.4.5.4); post-cloacal bones absent. Meatal closure muscle absent; anterior nuchal muscle originating mainly on second vertebra*; pectoralis muscles on each side crossing midline*, toes without complex internal musculature. Pupil approximately round with posteroventral quadrant of border less convex, or pupil vertically elliptical with un-notched borders; dorsal scaling consisting of small granules, nearly always without larger tubercles on body; preanal and femoral pores absent, cloacal tubercles and postcloacal sacs absent; toes simple without expanded adhesive pads beneath; tail strongly compressed laterally in males*, not or less so in females; in one species (P. celerrimus) an area of dark rugose scales in preanal region of males; flanks often with longitudinal series of short dark or reddish streaks separated by pale spots*. Largely diurnal and usually heliothermic; tail often raised and moved during intraspecific signalling*; voice little used [*Asterisks indicate features that are apparently largely or wholly confined to Pristurus among the Sphaerodactylidae. Gekkonidae and Phyllodactylidae; from ARNOLD 2009].

Diagnosis (gasperetti): A moderate sized species of Pristurus with a narrow head and somewhat laterally compressed body in males, where the neural spines form a distinct ridge along the back; transverse section of frontal bone concave above, its lateral downgrowths not meeting or fusing on mid-line or extending forwards onto nasal capsules; nasal process of premaxilla not extending backwards as far as the level of the anterior margins of the orbits, the posterior margin of its palatal section forming two gentle curves rather than a strong W-shape; clear and extensive contact between the palatal sections of the maxillae behind the premaxilla, postorbital bone relatively large; nearly always 23 presacral vertebrae with the lateral sections of the anterior zygapophyses folded right over the contiguous posterior zygapophyses of the next vertebra; three sternal and one complete xiphisternal rib; nostril situated be- tween the rostral, a supranasal and one or two postnasal scales; usually one (occasionally 2-4) scales between the two supranasals, upper palpebral fold narrow and scarcely projecting, subtibial scales enlarged; tail strongly compressed in males with a sagittal fringe of long scales above that does not extend on to body, and a similar but less developed fringe below. Most similar to Pristurusjlavipunctatus Riippell, 1835 but differs in the dorsal tail crest not extending on to the body, larger size (up to 38 mm from snout to vent compared with 33 mm in Arabian P. jlavipunctatus), more sombre colouring and stronger markings. Pristuruspopovi Arnold, 1982 has a broader head, a more extensive palpebral fold and less ventral tail cresting in males. Pristurus rupestris Blanford, 1874 is smaller, the downgrowths of the frontal bone meet and fuse beneath, the palpebral fold is better developed while tail fringing is often less so. In both these latter species, the transverse section of the frontal bone is not strongly concave above and the postorbital bone is smaller [ARNOLD 1986]. The common name of P. gasperetti is Wadi Kharrar Rock Gecko. 
EtymologyNamed after the yellow spots characteristic for this species. P. gasperetti was named after John Gasperetti (1920-2001), engineer and surveyor of various companies and governments in the Middle East and Field Associate of the Department of Herpetology of the California Academy of Sciences, San Francisco. 
References
  • Arnold E N 1986. New species of semaphore gecko (Pristurus: Gekkonidae) from Arabia and Socotra. Fauna of Saudi Arabia 8: 352-377
  • Arnold, E.N. 2009. Relationships, evolution and biogeography of Semaphore geckos, Pristurus (Squamata, Sphaerodactylidae) based on morphology. Zootaxa 2060: 1-21 - get paper here
  • Boulenger, G. A. 1896. A list of the reptiles and batrachians collected by Dr. Ragazzi in Shoa and Eritrea. Annali Mus. civ. Stor. nat. Genova, Giacomo Doria, (2) 16: 545-554 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • CUNNINGHAM, P. 2009. A contribution to the reptiles of the Farasan Islands, Saudi Arabia. African Herp News (50): 21-24
  • Duméril, A.M. C. and G. Bibron. 1836. Erpetologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 3. Libr. Encyclopédique Roret, Paris, 528 pp. - get paper here
  • Duméril, A.M.C., and G. Bibron. 1835. Erpétologie Générale ou Histoire Naturelle Complète des Reptiles, Vol. 2. Librairie Encyclopédique de Roret, Paris, iv + 680 p. - get paper here
  • Haas, Georg 1951. On the present state of our knowledge of the herpetofauna of Palestine. Bulletin of the Research Council of Israel 1: 67-95
  • Ineich, I. 1999. Reptiles & Amphibiens de la République de Djibouti. BNDB, UICN, PNUD, Djibouti & MNHN, Paris, 60 pp. - get paper here
  • Lanza, B. 1990. Amphibians and reptiles of the Somali Democratic Republic: check list and biogeography. Biogeographia, 14: 407-465 [1988]
  • Largen, M.J.; Spawls, S. 2006. Lizards of Ethiopia (Reptilia Sauria): an annotated checklist, bibliography, gazetteer and identification. Tropical Zoology 19 (1): 21-109 - get paper here
  • Largen, M.J.; Spawls, S. 2010. Amphibians and Reptiles of Ethiopia and Eritrea. Edition Chimaira, Frankfurt, 694 pp.
  • Loveridge,A. 1947. Revision of the African lizards of the family Gekkondiae. Bull. Mus. Comp. Zool. Harvard 98: 1-469 - get paper here
  • Marx,Hymen 1968. Checklist of the reptiles and amphibians of Egypt. Spec. Publ. U.S. Nav. Med. Res. Unit 3: 1-91
  • Mazuch, Tomáš 2013. Amphibians and Reptiles of Somaliland and Eastern Ethiopia. Tomáš Mazuch Publishing, 80 pp. ISBN: 978-80-905439-0-4 - get paper here
  • Milto, Konstantin D. 2017. New Records of Reptiles on the Red Sea Coast, Egypt, with Notes on Zoogeography Russ. J. Herpetol. 24 (1): 11-21 - get paper here
  • Parker, H. W. 1942. The lizards of British Somaliland. Bull. Mus. Comp. Zool. Harvard 91: 1—101 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Rösler, H.; Köhler, J. & Böhme, W. 2008. A new species of the diurnal gekkonid genus Pristurus Rüppell, 1835 from the Red Sea island Hanish al-Kabir, Yemen. Amphibia-Reptilia 29: 217-227 - get paper here
  • Rüppell, E. 1835. Neue Wirbelthiere zu der Fauna von Abyssinien gehörig, entdeckt und beschrieben. Amphibien. S. Schmerber, Frankfurt a. M. - get paper here
  • Schätti, B. & Gasperetti, J. 1994. A contribution to the herpetofauna of Southwest Arabia. Fauna of Saudi Arabia 14: 348-423
  • Scortecci, G. 1933. Descrizione preliminare di nuove specie e sottospecie de genere Pristurus della Somalia Italiana. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 72:242-244
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Sindaco, Roberto; Riccardo Nincheri, Benedetto Lanza 2014. Catalogue of Arabian reptiles in the collections of the “La Specola” Museum, Florence. Scripta Herpetologica. Studies on Amphibians and Reptiles in honour of Benedetto Lanza: pp. 137-164 - get paper here
  • van der Kooij, Jeroen 2001. The herpetofauna of the Sultanate of Oman: Part 2: the geckoes. Podarcis 1 (4): 105-120
 
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