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Proctoporus unsaacae DOAN & CASTOE, 2003

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Higher TaxaGymnophthalmidae, Sauria (lizards) 
Subspecies 
Common Names 
SynonymProctoporus unsaacae DOAN & CASTOE 2003
Proctoporus unsaacae — CASTOE et al. 2004 
DistributionPeru (Cusco)

Type locality: from Quello Uno (13° 21.887’ S, 71° 58.215’ W), a village near the town of Calca, Province of Calca, Department of Cusco, Peru; 3253 m elevation. Map legend:
Type locality - Type locality.
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
TypesHolotype: UTA R-51488 
CommentDiagnosis.—(1) Nasal divided, forming a pentagonal loreal scale; (2) median occipital present; (3) superciliaries four, first expanded onto dorsal surface of head; (4) palpebral eye-disc made up of a single, undivided scale; (5) genials two; (6) dorsal scales quadrangular, with low rounded keel, surrounded or not by longitudinal striations; (7) transverse rows of dorsals 36–42; (8) transverse ventral rows 18– 26; (9) a continuous series of small lateral scales separating dorsals and ventrals; (10) femoral pores per hind limb in males 5–7, in females 0–3; (11) preanal pores absent; (12) limbs not overlapping when adpressed against the body in adults; (13) a continuous series of dark-centered ocelli on the lateral surface; (14) venter cream, medial scales immaculate, ventrolateral scales with dark spot centered in each scale. Specimens of P. unsaacae show the pres-ence of an undivided palpebral eye-disc, which place them in the P. pachyurus group (Uzzell, 1970). Proctoporus unsaacae can be distin-guished from all other Proctoporus species except those in the P. pachyurus group by this character. Proctoporus unsaacae can be distinguished from P. bolivianus by a continuous series of lateral ocelli (in P. bolivianus, ocelli, if present, are faint and do not form a continuous series). It can be distinguished from P. guentheri by cream venter with ventrolateral scales that each have a dark spot (P. guentheri has yellow or orange venter without any spots). It can be distinguished from P. pachyurus by a lower number of transverse dorsal scale rows (47–61 in P. pachyurus; 36–42 in P. unsaacae). It can be distinguished from P. sucullucu by limbs not overlapping when adpressed. 
EtymologyEtymology: Named in honor of the herpetological research group at the Universidad Nacional de San Antonio Abad de Cusco, a university in Cusco, Peru, commonly referred to as UNSAAC. 
References
  • Castoe, T.A.; Doan, T.M. & Parkinson, C.L. 2004. Data partitions and complex models in Bayesian analysis: the phylogeny of Gymnophthalmid lizards. Systematic Biology 53 (3): 448-469 - get paper here
  • Doan, T. M. & Castoe, T.A. 2003. Using morphological and molecular evidence to infer species boundaries within Proctoporus bolivianus Werner (Squamata: Gymnopthalmidae). Herpetologica 59 (3): 432–449 - get paper here
  • Doan, T.F.; Castoe, T.A. & Arizábal Arriaga, W. 2005. Phylogenetic relationships of the genus Proctoporus sensu stricto (Squamata: Gymnophthalmidae), with a new species from Puno, Southeastern Peru. Herpetologica 61 (3): 325-336 - get paper here
  • Goicoechea, Noemí; José M. Padial, Juan C. Chaparro, Santiago Castroviejo-Fisher, Ignacio De la Riva 2013. Molecular phylogenetics, species diversity, and biogeography of the Andean lizards of the genus Proctoporus (Squamata: Gymnophthalmidae). Molecular Phylogenetics and Evolution, Volume 65, Issue 3, December 2012, Pages 953–964 - get paper here
  • Köhler, G. and E. Lehr 2004. Comments on Euspondylus and Proctoporus (Squamata: Gymnophthalmidae) from Peru, with the description of three new species and a key to the Peruvian species. Herpetologica 60 (4):501-518 - get paper here
 
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