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Psammodromus algirus (LINNAEUS, 1758)

IUCN Red List - Psammodromus algirus - Least Concern, LC

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Higher TaxaLacertidae (Gallotiinae), Sauria, Lacertoidea, Squamata (lizards) 
SubspeciesPsammodromus algirus algirus (LINNAEUS 1758)
Psammodromus algirus doriae BEDRIAGA 1886
Psammodromus algirus nollii FISCHER 1887
Psammodromus algirus ketamensis GALAN 1931 
Common NamesE: Algerian Psammodromus
G: Algerischer Sandläufer
D: Algerijnse Zandloper 
SynonymLacerta algira LINNAEUS 1758: 203
Tropidosaura algira — DUMÉRIL & BIBRON 1839: 168
Tropidosaura algira — BOETTGER 1874
Psammodromus algirus — BOULENGER 1887
Troidosaurus algira var. nolli FISCHER. 1887 (fide SCHLEICH et al. 1996)
Pantodactylus nicefori BURT & BURT 1931 (fide RUBIAL 1950)
Psammodromus algirus — ENGELMANN et al 1993
Psammodromus algirus — SCHLEICH, KÄSTLE & KABISCH 1996: 444
Psammodromus manuelae BUSACK, SALVADOR & LAWSON 2006
Psammodromus jeanneae BUSACK, SALVADOR & LAWSON 2006

Psammodromus algirus doriae BEDRIAGA 1886
Psammodromus algirus doriae — SINDACO & JEREMČENKO 2008

Psammodromus algirus nollii FISCHER 1887
Tropidosaura algira vor. Nollii FISCHER 1887
Psammodromus algirus nollii — SINDACO & JEREMČENKO 2008 
DistributionItaly (limited to the Isolotto dei Conigli near Lampedusa, Sicily) [E. RAZZETTI, pers. comm.]),
Portugal, Spain (incl. Mallorca), Andorra, Gibraltar, S France
N Tunisia, N Algeria, Morocco

Type locality: “Mauritania”

doriae: N Tunisia (Galita Archipelago)

ketamensis: N Morocco; type locality: “Telata de Ketama, Marruecos” [Morocco].

nollii: Morocco, Algeria (High Atlas)

manuelae: Spain (around Madrid); Type locality: Manzanares el Real, Madrid Province, Spain, 40°44' N, 3° 52' W.

jeanneae: Spain (Cádiz); Type locality: 25.6 km NE of Facinas (36° 08 N, 5° 42 W) along CA-221, Cádiz Province, Spain, elevation ca. 200 m. Map legend:
Type locality - Type locality.
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesType: BMNH 1946.8.4.69 (and possibly additional specimens).
Holotype: MNCN 7853. Paratypes: MNCN [ketamensis]
Holotype: MNCN 36792, an adult male, collected by J. Ibáñez, 24 June 1973 [manuelae]
Holotype: MNCN 11941 (S. D. Busack field series [SDB] 1526; GenBank DQ 150373 [NADH subunit 4] and DQ 150387 [NADH subunit 2]; MVZ 232055 [tissue sample]) [jeanneae]
Holotype: AMNH R1082; type locality: "Bogata, Colombia" (apparently in error) [Pantodactylus nicefori] 
CommentSubspecies after SCHLEICH, KÄSTLE & KABISCH 1996.

Synonymy: Psammodromus manuelae BUSACK, SALVADOR & LAWSON 2006 and Psammodromus jeanneae BUSACK, SALVADOR & LAWSON 2006 have been synonymized with P. algirus by Verdú-Ricoy et al. 2010.

Illustration of P.a. ketamensis in SCHLÜTER 2004.

Diagnosis (manuelae): One of two members in the Psammodromus algirus complex known to inhabit Europe. In addition to mtDNA and allozyme differences published earlier (Busack and Lawson in press), P. manuelae can be distinguished from the other known European member of the complex (see below) by, in males, having an average of eight (vs four) blue scales in the ocellus, and an ocellus area of approximately 4 mm² (vs 2 mm²). In addition, males tend to have a higher number of femoral pores (X = 18 vs X = 16) and dorsal scale rows (X [X meanin g average] = 25 vs X = 24) and, generally, a wider head (X = 11.7 mm vs 10.7 mm). Coloration in males differs from the other known European member of the complex in that the blue ocelli are bordered in black, as opposed to not being bordered in black, the dorsal surfaces of the head, body, limbs and tail are uniform brown as opposed to being uniform pale olive brown, the lateral band is dark brown with black scales and yellow spots tending to form vertical, and parallel, series that alternate along each side of the body, as opposed to being pale olive brown with very few black scales. Females tend to have a lower number of subdigital lamellae (X = 14 vs X = 15) and higher number of femoral pores (X = 16 vs X = 14) and dorsal scale rows (X = 25 vs X = 24), and, generally, a wider head (X = 10.2 mm vs 9.6 mm) than the other known European member of the complex. General coloration in female P. manuelae is lighter, but similar to that of males; females, however, have neither black scales nor yellow spots on the sides, and the small blue ocellus is bordered by brown instead of black.

Diagnosis (jeanneae): One of two members in the Psammodromus algirus complex known to inhabit Europe. In addition to mtDNA and allozyme differences published earlier (Busack and Lawson in press), P. jeanneae can be distinguished from P. manuelae by, in males, having an average of four (vs eight) blue scales in the ocellus, and an ocellus area of approximately 2 mm² (vs 4 mm²). In addition, males tend to have a lower number of femoral pores (X [X meaning average] = 16 vs X = 18) and dorsal scale rows (X = 24 vs X = 25), and, generally, a narrower head (X = 10.7 mm vs 11.7 mm). Coloration differs from P. manuelae in having dorsal surfaces of the head, body, limbs and tail uniform pale olivebrown as opposed to uniform brown, the two longitudinal stripes on each side of the body white as opposed to yellowish, and the lateral band between these stripes pale olive brown with very few black scales as opposed to dark brown with black scales and yellow spots. Females tend to have a higher number of subdigital lamellae (X = 15 vs X = 14) and lower number of femoral pores (X = 14 vs X = 16) and dorsal scale rows (X = 24 vs X = 25), and, generally, a narrower head (X = 9.6 mm vs 10.2 mm) than P. manuelae. Coloration in females is similar to that of males. 
EtymologyEtymology (manuelae): Named after Manuela González, wife of Alfredo Salvador.

Etymology (jeanneae). Named after Jeanne A. Visnaw, wife of Stephen D. Busack, who succumbed to ovarian cancer on 9 September 2005. Jeanne located and captured the first example of this species during 1982, and prepared both the voucher and its tissue for later examination. 
References
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