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Psammophis punctulatus DUMÉRIL, BIBRON & DUMÉRIL, 1854

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Higher TaxaPsammophiidae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Speckled Sand Racer, Sand snake 
SynonymPsammophis punctulatus DUMÉRIL, BIBRON & DUMÉRIL 1854: 897
Psammophis punctulatus — BOULENGER 1895
Psammophis punctulatus — SCORTECCI 1929: 305
Psammophis punctulatus — SCORTECCI 1939: 280
Psammophis punctulatus — LARGEN & SPAWLS 2010: 560
Psammophis punctulatus — WALLACH et al. 2014: 579
Psammophis punctulatus — SPAWLS et al. 2018: 438
Psammophis punctulatus — ŠMÍD et al. 2023 
DistributionS Egypt, Sudan (Jumhūriyyat), Republic of South Sudan (RSS), N Ethiopia, Eritrea, Djibouti, N Somalia

Type locality: “Taita”  
Reproductionoviparous 
TypesHolotype: MNHN-RA 6552 
DiagnosisAdditional details, e.g. a detailed description or comparisons (926 characters), are available for collaborators and contributors. Please contact us for details. 
CommentDistribution: CORKILL & COCHRANE 1966 list this species from the Arabian peninsula, probably in error. See map in Mazuch & Spawls 2023.

Subspecies: Psammophis punctulatus trivirgatus is now considered as a valid species. 
References
  • Baha El Din, Sherif and Osama Fathalla Ghazali 2005. On the occurrence of Psammophis punctulatus Bibron & Duméril 1854 in Egypt. Herpetological Bulletin 94: 4-5. - get paper here
  • Boulenger,G.A. 1895. Rettili e Batraci. in, Esplorazione del Giuba e dei suoi Affluenti compiuta dal Cap. V. Bottego durante gli Anni 1892-93 sotto gli auspicii della Società Geografica Italiana. Annali Mus. civ. Stor. nat. Giacomo Doria (2) 15: 9-18 - get paper here
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Cimatti, E. 2005. Scaly encounters in Kenya. Reptilia (GB) (42): 65-69 - get paper here
  • Corkill, N. L. and Cochrane, J. A. 1966. The snakes of the Arabian Peninsula and Socotra. J. Bombay nat. Hist. Soc. 62 (3): 475-506 (1965) - get paper here
  • Duméril, A. M. C., Bibron, G. & DUMÉRIL, A. H. A., 1854. Erpétologie générale ou histoire naturelle complète des reptiles. Tome septième. Deuxième partie, comprenant l'histoire des serpents venimeux. Paris, Librairie Encyclopédique de Roret: i-xii + 781-1536 - get paper here
  • Ineich, I. 1999. Reptiles & Amphibiens de la République de Djibouti. BNDB, UICN, PNUD, Djibouti & MNHN, Paris, 60 pp. - get paper here
  • Kirchhof S, Wasonga V, Mazuch T, Spawls S, Malonza KP. 2023. An annotated checklist of the herpetofauna of the Sibiloi National Park in northern Kenya based on field surveys. Amphibian & Reptile Conservation 17(1/2) [General Section]: 1–18 (e324) - get paper here
  • Kirchhof, S; Losogo, L B & Malonza, P K 2018. Psammophis punctulatus trivirgatus (Southern Speckled Sand Snake) Diet. Herpetological Review 49 (3): 552-553 - get paper here
  • Lanza, B. 1990. Amphibians and reptiles of the Somali Democratic Republic: check list and biogeography. Biogeographia, 14: 407-465 [1988] - get paper here
  • Largen, M.J.; Spawls, S. 2010. Amphibians and Reptiles of Ethiopia and Eritrea. Edition Chimaira, Frankfurt, 694 pp.
  • Largen,M.J. 1997. An annotated checklist of the amphibians and reptiles of Eritrea, with keys for their identification. Tropical Zoology 10: 63-115 - get paper here
  • Largen,M.J. & Rasmussen,J.B. 1993. Catalogue of the snakes of Ethiopia (Reptilia Serpentes), including identification keys. Tropical Zoology 6: 313-434 - get paper here
  • Massimo Capalbo 2024. The World's Most Dangerous Snakes - Volume I: From the fast-lightning Black Mamba to the insidious Hook-nosed Sea Snake, through the formidable Coastal Taipan. Privately published, 252 pp. - get paper here
  • Mazuch, Tomáš 2013. Amphibians and Reptiles of Somaliland and Eastern Ethiopia. Tomáš Mazuch Publishing, 80 pp. ISBN: 978-80-905439-0-4 - get paper here
  • Peters,W.C.H. 1878. Über die von Hrn. J. M. Hildebrandt während seiner letzten ostafrikanischen Reise gesammelten Säugethiere und Amphibien. Mber. Akad. Wiss. Berlin 1878: 194-209 - get paper here
  • Pitman,C.R.S. 1974. A guide to the snakes of Uganda. Codicote, Wheldon & Wesley, L., 290 pp.
  • Scortecci ,G. 1939. Spedizione zoologica del Marchese Saverio patrizi nel Basso Giuba e nell'Oltre Giuba. Giugno-agosto 1934. XII. Rettili Ofidi. Annali Mus. civ. Stor. nat. Giacomo Doria 58: 263-291 - get paper here
  • Scortecci, G. 1929. Rettili dell'Eritrea esistenti nelle Collezioni del Museo Civico de Milano. Atti della Società Italiana di Scienze Naturali, e del Museo Civico di Storia Naturale, Milano 67 (3-4): 290-339 [1928] - get paper here
  • Šmíd J, Fernández SM, Elmi HSA, Mazuch T 2023. Diversity of Sand Snakes (Psammophiidae, Psammophis) in the Horn of Africa, with the description of a new species from Somalia. Zoosystematics and Evolution 99(2): 345-361 - get paper here
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Spawls, Stephen; Tomáš Mazuch & Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • Trape, J.-F. 2023. Guide des serpents d’Afrique occidentale, centrale et d’Afrique du Nord. IRD Éditions, Marseille, 896 pp.
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Werner,F. 1919. Wissenschaftliche Ergebnisse der mit Unterstützung der Kaiserlichen Akademie der Wissenschaften in Wien aus der Erbschaft Treitl von F. Werner unternommenen zoologischen Expedition nach dem Anglo-Aegyptischen Sudan (Kordofan) 1914. IV. Bearbeitung de Denkschr. Akad. Wiss Wien, Math.-Naturw. Klasse 96: 437-509 - get paper here
 
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