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Psammophis aegyptius MARX, 1958

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Higher TaxaPsammophiidae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common Names 
SynonymPsammophis aegyptius MARX 1958: 194
Psammophis schokari aegyptius — KRAMER & SCHNURRENBERGER 1963
Psammophis aegyptius — WERNER 1985
Psammophis schokari aegyptius — BRANDSTÄTTER 1996
Psammophis aegyptius — SCHLEICH, KÄSTLE & KABISCH 1996: 513
Psammophis aegyptius —TRAPE & MANÉ 2006: 146
Psammophis aegyptius — WALLACH et al. 2014: 575
Psammophis aegyptius — BAR et al. 2021 
DistributionSE Algeria, S Egypt, Libya, Israel, Niger, Chad, Sudan (Jumhūriyyat)

Type locality: Siwa, Siwa Oasis, Western Desert Governorate, Egypt.  
Reproductionoviparous 
TypesHolotype: FMNH 75092. 
DiagnosisDiagnosis: A Psammophis with a stripe on the side of the head, running through the eye; belly finely punctulated; males with 188-199 ventrals and 17-19 scale rows at mid-body (from MARX 1958).


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CommentDistribution: For a map see Sindaco et al. 2013 and Bar et al. 2021. 
EtymologyNamed after its distribution in Egypt. 
References
  • ALOUFI, ABDULHADI A.; ZUHAIR S. AMR & MOHAMMAD A. ABU BAKER. 2022. New records of rare snakes from Jazan Province, Saudi Arabia (Reptilia: Serpentes). Zoology in the Middle East 0(0): 1–3. - get paper here
  • Bar, A., Haimovitch, G. and Meiri, S. 2021. Field guide to the amphibians and reptiles of Israel. Edition Chimaira, Frankfurt Am Main. ISBN 9783-89973-120-0 - get paper here
  • Bar, Aviad and Guy Haimovitch 2012. A Field Guide to Reptiles and Amphibians of Israel. Pazbar LTD, 246 pp. - get paper here
  • Bauer, Aaron M.; Jonathan C. DeBoer , Dylan J. Taylor 2017. Atlas of the Reptiles of Libya. Proc. Cal. Acad. Sci. 64 (8): 155-318 - get paper here
  • Brandstätter,F. 1996. Die Sandrennnattern. Die Neue Brehm-Bücherei 636, Magdeburg (Germany), 142 pp.
  • Chippaux, Jean-Philippe & Kate Jackson 2019. Snakes of Central and Western Africa. Johns Hopkins University Press, 448 pp. [detaileld review in HR 51 (1): 161] - get paper here
  • Geniez, Philippe 2018. Snakes of Europe, North Africa and the Middle East. Princeton University Press, 384 pp. - get paper here
  • Jestrzemski, Daniel 2018. Auf Schlangensuche im Heiligen Land – eine Woche Israel. Ophidia 12 (1): 2-17
  • Kark, Salit; Warburg, Ittai; Werner, Yehudah L 1997. Polymorphism in the snake Psammophis schokari on both sides of the desert edge in Israel and Sinai. Journal of Arid Environments 37 (3): 513-527 - get paper here
  • Kramer, E. & H. SCHNURRENBERGER 1963. Systematik, Verbreitung und Ökologie der Libyschen Schlangen. Rev. Sui. Zool. 70 (3): 453-568 - get paper here
  • Marx,H. 1958. Egyptian snakes of the genus Psammophis. Fieldiana: Zoology 39: 191-200 - get paper here
  • Meiri, Shai; Amos Belmaker, Daniel Berkowic, Kesem Kazes, Erez Maza, Guy Bar-Oz and Roi Dor 2019. A checklist of Israeli land vertebrates. Israel Journal of Ecology and Evolution - get paper here
  • Milto, Konstantin D. 2017. New Records of Reptiles on the Red Sea Coast, Egypt, with Notes on Zoogeography. Russ. J. Herpetol. 24 (1): 11-21 - get paper here
  • Milto, Konstantin D.; Samy A. Saber, Abdullah M. Nagy, Roman A. Nazarov, Daniel A. Melnikov, Natalia B. Ananjeva 2019. First Report on the Reptile Diversity of Wadi El Gemal National Park, Eastern Desert, Egypt. Russ. J. Herpetol. 26 (3): 175-184 - get paper here
  • Rouag R, Ziane N, De Sousa M 2024. A tentative list of reptilian fauna of Algeria and their conservation status. Biodiversity Data Journal 12: e120471 - get paper here
  • Schleich, H.H., Kästle,W., Kabisch, K. 1996. Amphibians and Reptiles of North Africa. Koeltz, Koenigstein, 627 pp.
  • Sindaco, R.; Alberto Venchi & Cristina Grieco 2013. The Reptiles of the Western Palearctic, Volume 2: Annotated Checklist and Distributional Atlas of the Snakes of Europe, North Africa, Middle East and Central Asia, with an Update to Volume 1. Edizioni Belvedere, Latina (Italy), 543 pp. - get paper here
  • Šmíd J, Fernández SM, Elmi HSA, Mazuch T 2023. Diversity of Sand Snakes (Psammophiidae, Psammophis) in the Horn of Africa, with the description of a new species from Somalia. Zoosystematics and Evolution 99(2): 345-361 - get paper here
  • Spawls, Stephen; Tomáš Mazuch & Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Trape J-F and Mané Y. 2015. The snakes of Niger. Amphibian & Reptile Conservation 9(2) [Special Section]: 39–55 (e110) - get paper here
  • Trape, J.-F. 2023. Guide des serpents d’Afrique occidentale, centrale et d’Afrique du Nord. IRD Éditions, Marseille, 896 pp.
  • Trape, J.-F. & Mané, Y. 2006. Guide des serpents d’Afrique occidentale. Savane et désert. [Senegal, Gambia, Mauritania, Mali, Burkina Faso, Niger]. IRD Editions, Paris, 226 pp. - get paper here
  • Vasconcelos-Gonçalves, Duarte; Fernando Martínez-Freiría, Pierre-André Crochet, Philippe Geniez, Salvador Carranza, José Carlos Brito 2017. The role of climatic cycles and trans-Saharan migration corridors in species diversification: Biogeography of Psammophis schokari group in North Africa. Molecular Phylogenetics and Evolution, 118: 64-74 - get paper here
  • Venchi, Alberto and Roberto Sindaco 2006. Annotated checklist of the reptiles of the Mediterranean countries, with keys to species identification. Part 2 -Snakes (Reptilia, Serpentes). Annali del Museo Civico di Storia Naturale "G. Doria", Genova, XCVIII: 259-364
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Werner Y L 1985. Defensive behaviour in a boigine snake: first record of throat inflation in Psammophis. ISRAEL JOURNAL OF ZOOLOGY 33(1-2) 1985: 69-71
  • Werner,Y.L. 1988. Herpetofaunal survey of Israel (1950-1985), with comments on Sinai and Jordan and on zoogeographical heterogeneity. in: Yom-Tov,Y. & Tchernov, E. (eds.) The zoogeography of Israel, Dr. W. Junk Publishers, Dordrecht (Netherlands); ISBN 90-6193-650-0
 
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