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Psammophis lineolatus (BRANDT, 1838)

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Higher TaxaPsammophiidae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Steppe Ribbon Racer
Chinese: 花条蛇 
SynonymColuber (Taphrometopon) lineolatus BRANDT 1838: 243
Coluber caspius LICHTENSTEIN 1823
Chorisodon Sibericum DUMERIL & BIBRON 1854: 902(fide PETERS 1861: 483)
Taphrometopon lineolatum — BOULENGER 1896: 151
Taphrometopon lineolatum — NIKOLSKY 1908: 343
Psammophis triticeus WALL 1912 (fide SMITH 1943)
Taphrometopon lineolatum — RENDAHL 1933: 30
Taphrometopon lineolatum — SCHMIDT 1939
Psammophis lineolatus — SMITH 1943: 367
Psammophis lineolatum — FRYNTA et al. 1997
Taphrometopon lineolatum — WALLACH et al. 2014: 695 
DistributionEast shore of Caspian Sea to S/E Kazakhstan, Azerbaijan, Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan, Afghanistan, Iran, Pakistan,
Mongolia, NW China (Gansu, Ningxia, Xinjiang)

Type locality: Transcaspia, also given as “M. Caspium” (Caspian Sea); restricted to the Mangystau Region of the Kazakhstan: Ungoza Mountain south of Sarytash Gulf, Mangystau (Mangyshlak) Penninsula (44°26´ N, 51°12´ E) by Doronin et al. 2020.  
Reproductionoviparous 
TypesHolotype: ZISP (ZIL) 2042, a 584 mm specimen (Karéline, 1837). 
DiagnosisDiagnosis: Nasal scale completely divided; Fourth, fifth and sixth upper labials touching eye; scale rows at midbody 15 – 16; ventrals 180; subcaudals 81; anal divided; carnivorous. (Baig et al. 2008)


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CommentSynonymy: see note in Zhao & Adler on the use of the name Coluber caspius. WALLACH et al 2014: 694 recognized Taphrometopon as valid genus (with 6 species) citing ZAHER et al. 2012 who actually do not mention the genus (although they do include some of its species in their phylogenetic tree).

Distribution: Has been erroneously (?) reported from India. For a map see Sindaco et al. 2013.

Type species: Coluber (Taphrometopon) lineolatus BRANDT 1836 is the type species of the genus Taphrometopon BRANDT 1838. 
References
  • Akhmedenov K. M., Bakiev A. G. 2021. The Steppe Ribbon Racer Psammophis lineolatus (Brandt, 1838) (Serpentes: Lamprophiidae) in Mangistau and Atyrau oblasts from the Republic of Kazakhstan. [in Russian] Proc. Inst. Zool., Kazakhstan, pp. 29-36 - get paper here
  • Anderson, S. C. & A. E. Leviton 1969. Amphibians and reptiles collected by the Street expedition to Afghanistan, 1965. Proc. Cal. Acad. Sci. (4) 37: 25-56 - get paper here
  • Baig, KJ, Masroor, R., and Arshad, M. 2008. Biodiversity and ecology of the herpetofauna of Cholistan Desert, Pakistan. Russ. J. Herpetol. 15 (3): 193-205 - get paper here
  • Boulenger, G.A. 1896. Catalogue of the snakes in the British Museum, Vol. 3. London (Taylor & Francis), xiv + 727 pp. - get paper here
  • Brandt, Johann F. 1838. Note sur quatre nouvelles especes de serpents de la cote occidentale de la mer Caspienne et la Perse septentrionale, decouvertes par M. Kareline. Bull. Acad. Sci. St. Petersburg 3: 241-244 [1836?] - get paper here
  • CHEN, M., LIU, J., CAI, B., LI, J., WU, N., & GUO, X. 2021. A new species of Psammophis (Serpentes: Psammophiidae) from the Turpan Basin in northwest China. Zootaxa 4974 (1): 116-134 - get paper here
  • CHEN, Minli; Jinlong LIU, Jun LI , Na WU, and Xianguang GUO 2021. Complete Mitochondrial Genome of the Steppe Ribbon Racer (Psammophis lineolatus): The First Representative from the Snake Family Psammophiidae and its Phylogenetic Implications. Asian Herpetological Research 12 (3): 295－307 - get paper here
  • CHIRIKOVA, M. A., Y. A. ZIMA, M. V. PESTOV, AND V. A. TERENTYEV 2020. Biodiversity of the Herpetofauna of the Muyunkum Desert, Kazakhstan. Herpetological Review 51: 438-446.
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