You are here » home advanced search Psammophis schokari

Psammophis schokari (FORSKÅL, 1775)

Can you confirm these amateur observations of Psammophis schokari?

Add your own observation of
Psammophis schokari »

Find more photos by Google images search: Google images

Higher TaxaPsammophiidae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Schokari Sand Racer, Forskal Sand Snake
G: Sandrennatter 
SynonymColuber schokari FORSKAL 1775: 14
Coluber lacrymans REUSS 1834: 139
Psammophis moniliger — DUMÉRIL & BIBRON 1854: 891
Psammophis punctatus DUMÉRIL & BIBRON 1854: 896
Psammophis sibilans var. hierosolimitana JAN 1863
Psammophis sindanus STOLICZKA 1872 (fide SMITH 1943)
Psammophis moniliger sp. var. hierosolymitana — BOETTGER 1879: 65
Psammophis sibilans var. punctata — BOETTGER 1885
Psammophis schokari BOULENGER 1896: 157
Psamophis schokari [sic] — SCHMIDT 1939
Psammophis sibilans schokari — LOVERIDGE 1940: 24
Psammophis schokari — SMITH 1943: 363
Psammophis schokari — HAAS 1957: 81
Psammophis schokari — LEVITON 1959: 457
Psammophis schokari schokari — GASPERETTI 1988: 265
Psammophis schokari — FRYNTA et al. 1997
Psammophis schokari — SCHLEICH, KÄSTLE & KABISCH 1996: 515
Psammophis schokari — SCHMIDT & KUNZ 2005
Psammophis schokari — WALLACH et al. 2014: 579
Psammophis schokari — BAR et al. 2021 
DistributionNW India (Rajasthan, Punjab, Jammu and Kashmir), Afghanistan (LEVITON 1959: 461), Pakistan, S Turkmenistan, Saudi Arabia, Qatar

Western Sahara ?, Morocco, Algeria, Libya, Egypt, Sinai, Israel, Mali, Niger ?, Mauritania, Nigeria, Sudan (Jumhūriyyat), Republic of South Sudan (RSS), Ethiopia, Eritrea, Somalia, Saudi Arabia (incl. Farasan Islands), Oman, United Arab Emirates (UAE), Kuwait, Syria, Jordan, Iraq, Iran (Kavir desert), Yemen

Type locality: Yemen; neotype locality: Hodeida, Yemen

hierosolimitana: Type locality: “Jérusalem” [Israel]  
Reproductionoviparous 
TypesNeotype: FMNH 66153, designated by Marx 1988 who apparently was not aware of a syntype in MCZ R-1974.
Holotype: ZSM 2122/0, adult [hierosolimitana] 
DiagnosisAdditional details, e.g. a detailed description or comparisons (1718 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSubspecies: Psammophis schokari aegyptius has been elevated to species status.

Synonymy: Loveridge 1940: 22

Distribution: not in Benin (pers. comm., L. Chirio, 23 Aug 2013). Not in Niger fide Trape & Mané 2015. See map in Burriel-Carranza et al. 2019 for map in UAE. For a map see Sindaco et al. 2013 and Bar et al. 2021. In Qatar fide Wikipedia.

Variation: Psammophis schokari (Colubridae) occurs in Israel and Sinai, Egypt, in three pholidotically indistinguishable morphs: (1) striped, with four dark longitudinal stripes; (2) non-striped, plain or lightly dotted; and (3) rear-striped, dark stripes on the posterior part merge anteriorly (this pattern is first described by KARK et al. 1997).

Habitat: partly arboreal (Harrington et al. 2018). 
EtymologyCorkill & Cochrane 1966 derive the name schokari from the Arabic 'shigari', which would mean 'from the trees'. However, some sources (such as Wikipedia) call it “Schokar's bronzeback”, indicating an eponym of unknown source. 
References
  • Adil, Saba; Shermeen Ijaz, Hira Aslam, Rimsha Kanwal and Sehrish Afsheen 2020. Diversity of amphibians and reptiles in Daphar Forest Sanctuary, district Mandi Bahauddin, Pakistan. Journal of Wildlife and Ecology 4(1):15-26
  • Al-Quran, S. 2009. The Herpetofauna of the Southern Jordan. American-Eurasian J. Agric. & Environ. Sci., 6 (4): 385-391 [this journal has a dubious record, see http://goo.gl/81SBPv]
  • Aloufi, A., Al Zayer, M., & Amr, Z. S. 2023. Reptiles and Amphibians along the Coastal Area of the Eastern Province, Saudi Arabia. Ecologica Montenegrina, 68, 66-87 - get paper here
  • Aloufi, Abdulhadi A.; Zuhair S. Amr, Mohammad A. Abu Baker 2021. Reptiles and Amphibians of Al Madinah Al Munawwarah Province, Saudi Arabia. Russian Journal of Herpetology 28 (3): 1 - get paper here
  • Alshammari, A. M. 2022. Four New Records of Snake Species in Ar'ar Region, Northern Border of Saudi Arabia. Jordan Journal of Natural History, 9(1), 41-48 - get paper here
  • Alshammari, A. M. and Busais, S. M. 2020. Distribution of snakes in Ha’il Province, Saudi Arabia, with an identification key to the species. Russian Journal of Herpetology 27 (1): 5-10. - get paper here
  • Alshammari, Ahmed Mohajja; Salem M. Busais, Abdel A. Ibrahim 2017. Snakes in the Province of Ha’il, Kingdom of Saudi Arabia, including two new records. Herpetozoa 30 (1/2): 59 - 63 - get paper here
  • Ananjeva, NB, Orlov NL; Khalikov RG; Darevsky IS; Ryabov IS & Barabanov AV 2006. The Reptiles of North Eurasia. Taxonomic Diversity, Distribution, Conservation Status [this comprises the territory of the former Soviet Union and Mongolia]. Pensoft Series Faunistica 47, 250 pp.
  • Anderson, John 1901. A list of the reptiles and batrachians obtained by Mr. A. Blayney Percival in southern Arabia. Proc. Zool. Soc. London 1901 (2): 137-152 - get paper here
  • Anderson, S.C. 1963. Amphibians and Reptiles from Iran. Proc. Cal. Acad. Sci. Ser. 4, 31(16): 417-498 - get paper here
  • Arnold E N 1980. The scientific results of the Oman flora and fauna survey 1977 (Dhofar). The reptiles and amphibians of Dhofar, southern Arabia. JOURNAL OF OMAN STUDIES SPECIAL REPORT (No. 2): 273-332 - get paper here
  • Bar, A., Haimovitch, G. and Meiri, S. 2021. Field guide to the amphibians and reptiles of Israel. Edition Chimaira, Frankfurt Am Main. ISBN 9783-89973-120-0 - get paper here
  • Bar, Aviad and Guy Haimovitch 2012. A Field Guide to Reptiles and Amphibians of Israel. Pazbar LTD, 246 pp. - get paper here
  • Barata, Mafalda; Ana Perera, D. James Harris, Arie van der Meijden, Salvador Carranza, Francisco Ceacero, Enrique García-Muñoz, Duarte Gonçalves, Sérgio Henriques, Fátima Jorge, Jonathan C. Marshall, Luis Pedrajas and Pedro Sousa 2011. New observations of amphibians and reptiles in Morocco, with a special emphasis on the eastern region. Herpetological Bulletin (116): 4 - get paper here
  • Bauer, Aaron M.; Jonathan C. DeBoer , Dylan J. Taylor 2017. Atlas of the Reptiles of Libya. Proc. Cal. Acad. Sci. 64 (8): 155-318 - get paper here
  • Boettger, O. 1879. Reptilien und Amphibien aus Syrien. Ber. Senck. Naturf. Ges. naturf. Ges. 1878-1879: 57-84 - get paper here
  • Boettger, O. 1885. Liste der von Hrn. Dr. med. W. Kobelt in Algerien und Tunisien gesammelten Kriechtiere. In W. Kobelt (editor), Reiseerinnerungen aus Algerien und Tunis. Berichte der Senckenbergischen Naturforschungs Gesellschaft 1885: 452–475 - get paper here
  • Boettger, O. [as O. Böttger] 1880. Die Reptilien und Amphibien von Syrien, Palaestina und Cypern. Ber. Senckenb. Naturforsch. Ges., Frankfurt/M., 1879-1880: 132-219 - get paper here
  • Boulenger, G.A. 1896. Catalogue of the snakes in the British Museum, Vol. 3. London (Taylor & Francis), xiv + 727 pp. - get paper here
  • BOURQUIN, O. 2011. The terrestrial reptiles of Sir Bani Yas Island, United Arab Emirates. African Herp News (53): 27-34 - get paper here
  • Brandstätter, F. 2002. Schlangen in der Bibel. Beiträge zur Literatur und Geschichte der Herpetologie und Terrarienkunde 2: 29-34
  • Brown, J.N.B. 1980. A New Snake From Suweihan Area. Bulletin of the Emirates Natural History Group (11 July 1980): http://www.enhg.org/bulletin/b11/11_29.htm
  • Burriel-Carranza B, Pedro Tarroso, Johannes Els, Andrew Gardner, Pritpal Soorae, Ahmed Ali Mohammed, Sai Ravi Krishna Tubati, Mohamed Mustafa Eltayeb, Junid Nazeer Shah, Héctor Tejero-Cicuéndez, Marc Simó-Riudalbas, Juan Manuel Pleguezuelos, Daniel F 2019. An integrative assessment of the diversity, phylogeny, distribution, and conservation of the terrestrial reptiles (Sauropsida, Squamata) of the United Arab Emirates. PLoS ONE 14(5): e0216273 - get paper here
  • Burriel-Carranza, B., Els, J., & Carranza, S. 2022. Reptiles & amphibians of the Hajar Mountains CSIC, Barcino Solucions Gràfiques, SL, Barcelona, - get paper here
  • Carranza S, Xipell M, Tarroso P, Gardner A, Arnold EN, Robinson MD, et al. 2018. Diversity, distribution and conservation of the terrestrial reptiles of Oman (Sauropsida, Squamata). PLoS One 13 (2): e0190389 - get paper here
  • Carranza, Salvador; Johannes Els; Bernat Burriel-Carranza 2021. A field guide to the reptiles of Oman. Madrid : Consejo Superior de Investigaciones Científicas, 223 pp. [review in HR 53 (3): 531] - get paper here
  • CHEN, M., LIU, J., CAI, B., LI, J., WU, N., & GUO, X. 2021. A new species of Psammophis (Serpentes: Psammophiidae) from the Turpan Basin in northwest China. Zootaxa 4974 (1): 116-134 - get paper here
  • Chippaux, Jean-Philippe & Kate Jackson 2019. Snakes of Central and Western Africa. Johns Hopkins University Press, 448 pp. [detaileld review in HR 51 (1): 161] - get paper here
  • Corkill, N. L. and Cochrane, J. A. 1966. The snakes of the Arabian Peninsula and Socotra. J. Bombay nat. Hist. Soc. 62 (3): 475-506 (1965) - get paper here
  • Cottone, A.M. & A.M. Bauer 2009. Sexual size dimorphism, diet, and reproductive biology of the Afro-Asian Sand Snake, Psammophis schokari (Psammophiidae) Amphibia-Reptilia, 30(3): 331-340. - get paper here
  • CUNNINGHAM, P. 2009. A contribution to the reptiles of the Farasan Islands, Saudi Arabia. African Herp News (50): 21-24 - get paper here
  • Disi, A.M.; Modry, D.; Nečas, P. & Rifai, L. 2001. Amphibians and reptiles of the Hashemite Kingdom of Jordan. Edition Chimaira, Frankfurt, 408 pp.
  • Duméril, A. M. C., Bibron, G. & DUMÉRIL, A. H. A., 1854. Erpétologie générale ou histoire naturelle complète des reptiles. Tome septième. Deuxième partie, comprenant l'histoire des serpents venimeux. Paris, Librairie Encyclopédique de Roret: i-xii + 781-1536 - get paper here
  • Dunaev E.A., Orlova V.F. 2003. Diversity of snakes (on the materials of the exposition of the Zoological Museum of the Moscow State University). Moscow: Moscow State University Press, 376 pp
  • Ebrahimipour, Farnaz; Eskandar Rastegar-Pouyani, Batoul Ghorbani 2016. A Preliminary Study of the Reptile’s Fauna in Northwestern Yazd Province, Iran. Russ. J. Herpetol. 23 (4): 243-248 - get paper here
  • Egan, D. 2007. Snakes of Arabia. Motivate Publishing, Dubai, 208 pp.
  • Eissa, S.M. & EL ASSY, Y.S. 1975. Record of certain reptilian species found in Kuwait. Jour. Univ. Kuwait (Sci.), 2: 123-146 - get paper here
  • Elbahi A, Lawton C, Oubrou W, El Bekkay M, Hermas J, Dugon M 2022. Reptile biodiversity in Souss-Massa National Park: an internationally important hotspot in the Mediterranean region. Biodiversity Data Journal 10: e79088 - get paper here
  • Fathnia, B.; N. Rastegar Pouyani; H. Darvishnia; M. Rajabzadeh 2010. The snake fauna of Ilam Province, southwestern Iran. Iranian Journal of Animal Biosystematics 6 (1) - get paper here
  • Forskål,P. 1775. Descriptiones animalium, avium, amphibiorum, piscium, insectorum, vermium; quae in itinere Orientali observavit Petrus Forskål. Mölleri, Hauniae, xxxiv + 164 pp. - get paper here
  • Frynta,D. et al. 1997. Results of the Czech biological expedition to Iran. Part 1. Notes on the distribution of amphibians and reptiles. Acta Soc. Zool. Bohem. 61: 3-17
  • Gardner, A.S. 2013. The amphibians and reptiles of Oman and the UAE. Edition Chimaira, 480 pp.
  • Gasperetti,J. 1988. Snakes of Arabia. Fauna of Saudi Arabia 9: 169-450
  • Geniez, P.; Mateo, J.A.; Geniez, M. & Pether, J. 2004. The amphibians and reptiles of the Western Sahara (former Spanish Sahara) and adjacent regions. Edition Chimaira, Frankfurt, 228 pp. [reviewed in Reptilia GB 44: 81]
  • Geniez, Philippe 2018. Snakes of Europe, North Africa and the Middle East. Princeton University Press, 384 pp. - get paper here
  • Goldberg, STEPHEN R. 2015. Reproduction in the Schokari sand racer, Psammophis schokari (Squamata: Colubridae) from Israel. Herpetological Bulletin (131) - get paper here
  • Goldberg, Stephen R. and Charles R. Bursey. 2015. Psammophis schokari (Schokari sand racer) endoparasite. Herpetological Review 46 (4): 648-649 - get paper here
  • Grossmann, W., Kowalski, T., Zwanzig, B. M., Zilger, H. J. and Gebhart, J. 2020. Herpetological observations in Oman: Al Mughsayl and Wadi Ashawq in Dhofar, Part 1. Amphibia: Anura; Reptilia: Squamata: Sauria. Sauria 42 (1): 1-20.
  • Gruber, U. 2009. Die Schlangen Europas, 2. Aufl. Kosmos Naturführer, 266 pp.
  • Haas G. and Werner, Y. L. 1969. Lizards and snakes from Southwestern Asia, collected by Henry Field. Bull. Mus. Comp. Zool. Harvard 138: 327-406. - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Hellmich, W. 1959. Bemerkungen zu einer kleinen Sammlung von Amphibien und Reptilien aus Süd-Persien. Opuscula Zoologica 35: 1-9 - get paper here
  • Ibrahim, Adel A. 2013. The Herpetology of the Suez Canal Zone, Egypt. Vertebrate Zoology 63 (1): 87–110 - get paper here
  • Jablonski D, Regan JM, Holzheuser C, Farooqi J, Basit A, Masroor R 2019. Additional data to the herpetofauna of Afghanistan. Herpetozoa 32: 177-193 - get paper here
  • Jongbloed, M. 2000. Field Guide to the reptiles and amphibians of the UAE - Wild about reptiles. Barkers Trident Communications, 116 pp.
  • Kamali, Kamran 2020. A Field Guide to the Reptiles and Amphibians of Iran. Edition Chimaira, Frankfurt am Main, Germany (www.chimaira.de). 574 pp.
  • Kramer, E. & H. SCHNURRENBERGER 1963. Systematik, Verbreitung und Ökologie der Libyschen Schlangen. Rev. Sui. Zool. 70 (3): 453-568 - get paper here
  • Lanza, B. 1990. Amphibians and reptiles of the Somali Democratic Republic: check list and biogeography. Biogeographia, 14: 407-465 [1988] - get paper here
  • Largen, M.J.; Spawls, S. 2010. Amphibians and Reptiles of Ethiopia and Eritrea. Edition Chimaira, Frankfurt, 694 pp.
  • Largen,M.J. & Rasmussen,J.B. 1993. Catalogue of the snakes of Ethiopia (Reptilia Serpentes), including identification keys. Tropical Zoology 6: 313-434 - get paper here
  • Laurent, P. 1935. Contribution à la connalssance de la faune des Vertebres du Maroc (Batraciens, Reptiles, Mammiferes). Bull. Soc. d’Hist. nat. de l’Afrique du Nord 26: 344-359
  • Leviton, A.E. 1959. Systematics and Zoogeography of Philippine Snakes. Unpublished PhD Thesis.
  • Leviton, Alan E. 1959. Report on a collection of reptiles from Afghanistan. Proc. Cal. Acad. Sci. 29 (12): 445-463 - get paper here
  • Leviton,A.E. & Anderson,S.C. 1967. Survey of the reptiles of the Sheikdom of Abu Dhabi, Arabian Peninsula. Part II. Systematic account of the collction of reptiles made in the Sheikdom of Abu Daby by John Gasperetti. Proc. Cal. Acad. Sci. (4) 35: 157-192 - get paper here
  • Martínez del Mármol, Gabriel; D. James Harris, Philippe Geniez, Philip de Pous, and Daniele Salvi 2019. Amphibians and Reptiles of Morocco. Edition Chimaira, Frankfurt am Main, Germany, 478 pp - get paper here
  • Marx, H. 1988. The colubrid snake, Psammophis schokari, from the Arabian Peninsula. Fieldiana Zool. New Ser. 40 (1383): 1-16 - get paper here
  • Marx,H. 1958. Egyptian snakes of the genus Psammophis. Fieldiana: Zoology 39: 191-200 - get paper here
  • Masroor, R. 2012. A Contribution to the Herpetology of Northern Pakistan. SSAR, Ithaca [review in JoTT 4(6): 2670] - get paper here
  • Mazuch, Tomáš 2013. Amphibians and Reptiles of Somaliland and Eastern Ethiopia. Tomáš Mazuch Publishing, 80 pp. ISBN: 978-80-905439-0-4 - get paper here
  • Mármol, Gabriel Martínez del; Ander Izagirre-Egaña, Sonke Frahm & Marco Sassoè 2023. Herpetofauna of Mauritania: results of a field survey. Bol. Asoc. Herpetol. Esp. (2023) 34 Preprint-101 - get paper here
  • Mediani, Mohamed; José Carlos Brito, Soumia Fahd 2015. Atlas of the amphibians and reptiles of northern Morocco: updated distribution and patterns of habitat selection. Basic and Applied Herpetology 29: - get paper here
  • Meiri, Shai; Amos Belmaker, Daniel Berkowic, Kesem Kazes, Erez Maza, Guy Bar-Oz and Roi Dor 2019. A checklist of Israeli land vertebrates. Israel Journal of Ecology and Evolution - get paper here
  • Mertens, R. 1922. Reptilien aus Palästina - Nachtrag. Zool. Anz. 1/2 (54): 47-47
  • Mouane, A., Harrouchi, A. K., Ghennoum, I., Sekour, M., & Chenchouni, H. 2024. Amphibian and reptile diversity in natural landscapes and human-modified habitats of the Sahara Desert of Algeria: A better understanding of biodiversity to improve conservation. Elementa: Science of the Anthropocene, 12(1) - get paper here
  • Murthy, T.S.N. 2010. The reptile fauna of India. B.R. Publishing, New Delhi, 332 pp.
  • Müller, J. 1996. Angaben über einige Amphibien - und Reptilienfunde in Jordanien. Sauria 18 (3): 3-10 - get paper here
  • Nilson,G. & Andren,C. 1981. Die Herpetofauna des Kavir-Schutzgebietes, Kavir-Wüste, Iran. Salamandra 17 (3-4): 130-146 - get paper here
  • Osman, H.E.S.I., & Sir, N.T.E. 1988. The snakes of the Sudan. 1. The snakes of Khartoum Province. The Snake 20 (1): 74-79.
  • Oudjani, David, Michel Aymerich and Mohamed Mediani. 2016. Rhagerhis moilensis (REUSS, 1834), as prey of Psammophis schokari (FORSKÅL, 1775). Herpetozoa 28 (3/4): 184-187 - get paper here
  • O’Shea, M. 2018. The Book of Snakes. Ivy Press / Quarto Publishing, London, - get paper here
  • Padial, J. M. 2006. COMMENTED DISTRIBUTIONAL LIST OF THE REPTILES OF MAURITANIA (WEST AFRICA). Graellsia, 62(2): 159-178 - get paper here
  • Rajabizadeh, M. 2018. Snakes of Iran. [in Farsi] Tehran: Iranshenasi, 496 pp. [2017 ?] - get paper here
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Rato, C; Brito, JC; Carretero, MA; Larbes, S; Shacham, B; Harris, DJ 2007. Phylogeography and genetic diversity of Psammophis schokari (Serpentes) in North Africa based on mitochondrial DNA sequences. African Zoology 42 (1): 112-117 - get paper here
  • Reuss, A. 1834. Zoologische Miscellen. Reptilien, Ophidier. [Coluber albiventris, Echis pavo]. Mus. Senckenbergiana, Frankfurt/M., 1: 129-162. - get paper here
  • Rhadi, Fadhel Aabbas;Rihab Ghaleb Mohammed, Nasrullah Rastegar-Pouyani,Eskandar Rastegar-Pouyani, Seyyed Saeed Hosseinian Yousefkhani 2017. On the Snake Fauna of Central and Southern Iraq and Some Zoogeographic Remarks. Russ. J. Herpetol. 24 (4): 251-266 - get paper here
  • Rouag R, Ziane N, De Sousa M 2024. A tentative list of reptilian fauna of Algeria and their conservation status. Biodiversity Data Journal 12: e120471 - get paper here
  • Sahi D.N., Koul S. 2020. Annotated List of Amphibians and Reptiles of Jammu and Kashmir State. In: Dar G. & Khuroo A. (eds) Biodiversity of the Himalaya: Jammu and Kashmir State. Topics in Biodiversity and Conservation, vol 18. Springer, Singapore - get paper here
  • Schätti, B. & Gasperetti, J. 1994. A contribution to the herpetofauna of Southwest Arabia. Fauna of Saudi Arabia 14: 348-423
  • Schleich, H.H. 1987. Contributions to the herpetology of Kouf National Park (NE-Libya) and adjacent areas. Spixiana 10 (1): 37-80. - get paper here
  • Schleich, H.H., Kästle,W., Kabisch, K. 1996. Amphibians and Reptiles of North Africa. Koeltz, Koenigstein, 627 pp.
  • Schlüter, U. 2006. Sandliebhaber - die psammophilen Nattern Nordafrikas. Reptilia (Münster) 11 (60): 72-80 - get paper here
  • Schmidt, D. & Kunz, K. 2005. Ernährung von Schlangen. Natur und Tier Verlag, Münster, 159 pp. - get paper here
  • Schmidt, K. P. & Marx, H. 1956. The herpetology of Sinai. Fieldiana 39 (4): 21-40 - get paper here
  • Schmidt,K.P. 1939. Reptiles and amphibians from Southwestern Asia. Publ. Field Mus. nat. Hist., zool. Ser., 24: 49-92 - get paper here
  • Schnurrenberger, Hans 1963. Fishes, amphibians, and reptiles of two Libyan oases. Herpetologica 18 (4): 270-273 - get paper here
  • Scortecci, G. 1932. Rettili dello Yemen. Atti soc. Ital. Sci.Nat. 71: 39-49 - get paper here
  • Sharma, R. C. 2004. Handbook Indian Snakes. AKHIL BOOKS, New Delhi, 292 pp.
  • Shehzad, M., Ullah, F., Khan, S. N., Majid, A., Rais, M., Khan, M. A., ... & Khan, S. 2023. Squamate Fauna of Southern Khyber Pakhtunkhwa (Tank and Lakki Marwat Districts), Pakistan. Pakistan J. Zool. 2023: 1-7 - get paper here
  • Sindaco, R.; Alberto Venchi & Cristina Grieco 2013. The Reptiles of the Western Palearctic, Volume 2: Annotated Checklist and Distributional Atlas of the Snakes of Europe, North Africa, Middle East and Central Asia, with an Update to Volume 1. Edizioni Belvedere, Latina (Italy), 543 pp. - get paper here
  • Sindaco, Roberto; Riccardo Nincheri, Benedetto Lanza 2014. Catalogue of Arabian reptiles in the collections of the “La Specola” Museum, Florence. Scripta Herpetologica. Studies on Amphibians and Reptiles in honour of Benedetto Lanza: pp. 137-164 - get paper here
  • Singh J, Dutta S.K., Singh H 2021. Herpetofauna of Punjab: The Field Guide. New Era Book Agency, Chandigarh, 82 pp.
  • Šmíd J, Fernández SM, Elmi HSA, Mazuch T 2023. Diversity of Sand Snakes (Psammophiidae, Psammophis) in the Horn of Africa, with the description of a new species from Somalia. Zoosystematics and Evolution 99(2): 345-361 - get paper here
  • Smith, M.A. 1943. The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp.
  • Sow, Andack Saad; Duarte Vasconcelos Gonçalves, Fábio Vieira Sousa, Fernando Martínez- Freiría, Frederico Santarém, Guillermo Velo-Antón, Hamidou Dieng, João Carlos Campos, Saer Khayar Diagne, Zbyszek Boratyński, José Carlos Brito 2017. Atlas of the distribution of amphibians and reptiles in the Diawling National Park, Mauritania. Basic and Applied Herpetology - get paper here
  • Sow, Andack Saad; Fernando Martínez-Freiría, Pierre-André Crochet, Philippe Geniez, Ivan Ineich, Hamidou Dieng, Soumia Fahd, José Carlos Brito 2015. Atlas of the distribution of reptiles in the Parc National du Banc d’Arguin, Mauritania. Basic and Applied Herpetology - get paper here
  • Spawls, Stephen; Tomáš Mazuch & Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Steindachner, Franz 1903. Batrachier und Reptilien von Südarabien und Sokotra, gesammelt während der südarabischen Expedition der kaiserlichen Akademie der Wissenschaften. Sitzungsb. Akad. Wiss. Wien, math.-naturwiss. Kl., 112, Abt. 1: 7-14 - get paper here
  • Trape, J.-F. 2023. Guide des serpents d’Afrique occidentale, centrale et d’Afrique du Nord. IRD Éditions, Marseille, 896 pp.
  • Trape, J.-F. & Mané, Y. 2006. Guide des serpents d’Afrique occidentale. Savane et désert. [Senegal, Gambia, Mauritania, Mali, Burkina Faso, Niger]. IRD Editions, Paris, 226 pp. - get paper here
  • Trape, Jean-François & Youssouph Mané 2017. The snakes of Mali. Bonn zoological Bulletin 66 (2): 107–133 - get paper here
  • Trapp, B. 2006. Dünne Luft - eine Exkursion ins Hochland des Irans und die Türkei. Reptilia (Münster) 11 (59): 40-49 - get paper here
  • Ullenbruch, K.; Grell, O.; Böhme, W. 2010. Reptiles from southern Benin, West Africa, with the description of a new Hemidactylus (Gekkonidae), and a country-wide checklist. Bonn Zool. Bull. 57 (1): 31-54 - get paper here
  • van der Kooij, Jeroen 2001. The herpetofauna of the Sultanate of Oman: Part 4: The terrestrial snakes. Podarcis 2 (2): 54-65 - get paper here
  • Vasconcelos-Gonçalves, Duarte; Fernando Martínez-Freiría, Pierre-André Crochet, Philippe Geniez, Salvador Carranza, José Carlos Brito 2017. The role of climatic cycles and trans-Saharan migration corridors in species diversification: Biogeography of Psammophis schokari group in North Africa. Molecular Phylogenetics and Evolution, 118: 64-74 - get paper here
  • Venchi, Alberto and Roberto Sindaco 2006. Annotated checklist of the reptiles of the Mediterranean countries, with keys to species identification. Part 2 -Snakes (Reptilia, Serpentes). Annali del Museo Civico di Storia Naturale "G. Doria", Genova, XCVIII: 259-364
  • Wachtel, Eric 2019. Marokko: Auf Schlangensuche im Nordwesten Afrikas. Ophidia 13 (1): 3-17
  • Wall, F. 1911. Remarks on a snake collection in the Quetta museum. J. Bombay Nat. Hist. Soc. 20: 1033-1042 - get paper here
  • Wall,F. 1908. Notes on a collection of snakes from Persia. J. Bombay nat. Hist. Soc. 18: 795-805 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Werner, F. 1897. Über Reptilien und Batrachier aus Togoland, Kamerun und Tunis aus dem Kgl. Museum für Naturkunde in Berlin. Verh. Zool.-Bot. Ges., Wien 47: 395-407 - get paper here
 
External links  
Is it interesting? Share with others:


Please submit feedback about this entry to the curator