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Pseudocalotes ziegleri HALLERMANN, TRUONG, ORLOV & ANANJEVA, 2010

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Ziegler’s Tree Lizard
Vietnamese: Nhong zig-lo 
SynonymPseudocalotes ziegleri HALLERMANN, TRUONG, ORLOV & ANANJEVA 2010
Pseudocalotes ziegleri — BÖHME 2014: 127 
DistributionC Vietnam (Kon Tum)

Type locality: Nuoc Ka forest, near Mang Canh, Kon Plong District, Kon Tum Province, Central Vietnam (14°41’ 57’’ N 108°12’ 5.7’’ E at an elevation of approximately 1200 m.  
Reproduction 
TypesHolotype: IEBR 330, an adult male collected on 10 January 2001 by Nguyen Quang Truong. Paratypes: ZFMK 88832 (IEBR A.0834), juv., from Mang Canh, Kon Plong District, Kon Turn Province, Vietnam (14°41'N, 108° 19' E, 1150-1250 m a.s.I.), collected by N. ORLOV, Ho, T.C., and R. NAZAROV, May 1June 2006; ZISP, IEBR. 
DiagnosisDiagnosis: Pseudocalotes ziegleri sp. nov. differs from Pseudocalotes brevipes and P. microlepis in having fewer scales around midbody 57–64 (mean 60.2, n = 5) vs. 69 – 78 (mean 72.4, n = 17) in P. brevipes and 65 – 72 (mean 69.7, n = 6) in P. microlepis. The new species is similar to P. brevipes in having the same character of subdigital lamellae on third toe, however, it further dif fers from the latter by lacking white spots on the elbows, by having lateral scales rhomboidal and smooth (vs. triangular and feebly keeled in P. brevipes), and a relatively larger tympanum (ratio of orbit/tympanum 1.84 vs. 2.23 in P. brevipes) (Fig. 4). Whereas P. microlepis has a homogenous body scalation with nearly cycloid lateral body scales, with only the posterior point of the scale overlapping (Fig. 4). The new species somewhat resembles P. poilani in midbody scales rows (56 – 60 rows, mean 58.3, n = 3) and P. floweri (51 – 61 rows, mean 55.7, n = 4). However, it can be distinguished from P. floweri and P. poilani by having the leading edges of subdigital scales under third-toe triangular and pointed (vs. with blade-like lateral extensions in both latter species) (Fig. 5). P. poilani differs from both P. ziegleri sp. nov. and P. floweri by having a transverse gular fold in all males, females and juveniles and a heart-like black marking in males (vs. longitudinal gular fold or no fold in congeners, Fig. 6). P. ziegleri sp. nov. is distinguishable from P. flavigula by having more scales around midbody (57 – 64 vs. 38 – 40). The new species differs from Pseudocalotes dringi, P. khaonanensis, P. saravacensis, and P. tympanistriga in having the leading edges of the subdigital scales under of third toe modified to a triangular and pointed confition (vs. unmodified in remaining species). P. ziegleri sp. nov. differs from P. larutensis by having five or more scales on canthus rostralis between nasal and first supraorbital (vs. four in P. larutensis). 
Comment 
EtymologyThe new species is named after Dr. Thomas Ziegler (Cologne Zoo), to honor his remarkable contributions to extensive research on Vietnamese herpetology. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Böhme, Wolfgang 2014. Herpetology in Bonn. Mertensiella 21. vi + 256 pp. - get paper here
  • Hallermann, Jakob; Nguyen Quang Truong, Nikolai Orlov, and Natalia Ananjeva 2010. A new species of the genus Pseudocalotes (Squamata: Agamidae) from Vietnam. Russ. J. Herpetol. 17 (1): 31-40 - get paper here
  • Milto, Konstantin D.; Andrei V. Barabanov 2012. A Catalogue of the Agamid and Chamaeleonid Types in the Collection of the Zoological Institute, Russian Academy of Sciences, St. Petersburg. Russ. J. Herpetol. 19 (2): 155-170 - get paper here
  • Nguyen, Thomas Ziegler & Truong Quang; 2010. New discoveries of amphibians and reptiles from Vietnam. Bonn Zool. Bull. 57 (2): 137–147 - get paper here
  • Xu, Y., Gong, Y., Hou, M., Weng, S., Liu, S., Deng, J., ... & Peng, L. 2024. A New Species of the Genus Pseudocalotes (Squamata: Agamidae) from Southwest Yunnan, China. Animals, 14(6): 826 - get paper here
 
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