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Pseudocophotis kontumensis ANANJEVA, ORLOV, TRUONG & NAZAROV, 2007

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common Names 
SynonymPseudocophotis kontumensis ANANJEVA, ORLOV, TRUONG & NAZAROV 2007
Pseudocophotis kontumensis — MILTO & BARABANOV 2012: 166 
DistributionC Vietnam (Kon Tum Province)

Type locality: Kon Du, Mang Canh village, Konplong
district, Kon Tum Province, Vietnam (14°41’  
Reproductionoviparous 
TypesHolotype: ZISP 24251, adult male, collected on April 7, 2005, by Nikolai L. Orlov, Sergei Ryabov, and Ho Thu Cuc. 
DiagnosisDiagnosis: Pseudocophotis kontumensis differs from other agamid lizards known to occur in Vietnam based on morphological characters. It is a middle-sized agamid lizard with laterally compressed body, very large dorsal and lateral scales, smaller keeled ventral scales, hidden tympanum, fold across the throat and relatively short flexible prehensile tail. No gular sac. From Acanthosaura P. kontumensis can be distinguished by the absence of postorbital and nuchal spines; from Calotes — by heterogeneous unequal dorsal scales intermixed with larger irregular scales. From both genera it differs by tympanum covered with skin. From Bronchocela it is distinguished by its much shorter tail. From Gonocephalus P. kontumensis differs by hidden tympanum, much smaller adult size and smaller ventral scales in comparison with dorsal.
P. kontumensis differs from 3 species of Pseudocalotes occurring in Vietnam (Ananjeva et al., 2007; Bain et al., 2007): P. brevipes, P. microlepis, and P. floweri by hidden tympanum and smaller scale number around the midbody. P. brevipes and P. microlepis have much shorter tail (Table 1; Hallermann and McQuire, 2001: Table 1). Japalura species with hidden tympanum (including Vietnamese J. chapaense and J. fasciata) have much longer tail and limbs (Table 1; Ota, 1989: Tables 1 and 2). Pseudocophotis kontumensis differs from P. sumatrana by higher number of scales around midbody [from ANANJEVA et al. 2007]. 
CommentAbundance: only known from its original description (Meiri et al. 2017). This is one of the species called 'lost' and 'rediscovered' by Lindken et al. 2024.

Synonymy: Denzer et al. 2021 question the placement of kontumensis in Pseudocophotis, but given the lack of molecular data there is currently no better solution. 
Etymologynamed after the type locality. 
References
  • Aguilar-López JL, Luría-Manzano R, Pineda E, Canseco-Márquez L 2021. Selva Zoque, Mexico: an important Mesoamerican tropical region for reptile species diversity and conservation. ZooKeys 1054: 127-153 - get paper here
  • Ananjeva, Natalia B;Nikolai L. Orlov, Quang Truong Nguyen and Roman A. Nazarov. 2007. A new species of Pseudocophotis (Agamidae: Acrodonta: Lacertilia: Reptilia) from central Vietnam. Russ. J. Herpetol. 14 (2):153-160 - get paper here
  • Bain, Raoul H.; Truong, Nguyen Quang; van Kien, Doan 2007. New herpetofaunal records from Vietnam. Herpetological Review 38 (1): 107-117 - get paper here
  • Bobrov V.V., Semenov D.V. 2008. Lizards of Vietnam [in Russian]. Moscow, 236 pp.
  • Denzer, Wolfgang; Esther Dondorp, Thore Koppetsch, Karien Lahaise, Ulrich Manthey, Maria Mostadius & Wolfgang Böhme 2021. Nomenclatural and morphological notes on the rare agamid lizard Pseudocophotis sumatrana (Hubrecht, 1879) (Squamata: Agamidae: Draconinae). RAFFLES BULLETIN OF ZOOLOGY 69: 448–462 - get paper here
  • Hallermann, Jakob and Jimmy A. McGuire 2001. A new species of Pseudocalotes (Squamata: Agamidae) from Bukit Larut, West Malaysia. Herpetologica 57 (3):255-265 - get paper here
  • Lindken T.; Anderson, C. V., Ariano-Sánchez, D., Barki, G., Biggs, C., Bowles, P., Chaitanya, R., Cronin, D. T., Jähnig, S. C., Jeschke, J. M., Kennerley, R. J., Lacher, T. E. Jr., Luedtke, J. A., Liu, C., Long, B., Mallon, D., Martin, G. M., Meiri, 2024. What factors influence the rediscovery of lost tetrapod species? Global Change Biology, 30: 1-18 - get paper here
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Milto, Konstantin D.; Andrei V. Barabanov 2012. A Catalogue of the Agamid and Chamaeleonid Types in the Collection of the Zoological Institute, Russian Academy of Sciences, St. Petersburg. Russ. J. Herpetol. 19 (2): 155-170 - get paper here
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Ota,H. 1989. A new species of Japalura (Agamidae: Lacertilia:Reptilia) from Taiwan. Copeia 1989 (3): 569-576 - get paper here
 
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