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Pseudonaja mengdeni WELLS & WELLINGTON, 1985

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Higher TaxaElapidae (Hydrophiinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common NamesE: Gwardar, Western brownsnake 
SynonymPseudonaja mengdeni WELLS & WELLINGTON 1985
Pseudonaja kellyi WELLS & WELLINGTON 1985
Pseudonaja kellyi — WELLS 2002
Pseudonaja mengdeni — WELLS 2002
Pseudonaja mengdeni — WELLS 2002
Pseudonaja mengdeni — SKINNER 2009
Pseudonaja mengdeni — WALLACH et al. 2014: 605
Pseudonaja mengdeni — MIRTSCHIN et al. 2017 
DistributionAustralia (Western and central Australia, from Carnarvon, Western Australia, to Mootwingee National Park, western New South Wales)

Type locality: 2 km east of Maryvale, Northern Territory.  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: NTM R1989 
DiagnosisDiagnosis: “A member of the Pseudonaja nuchalis complex, readily identified by the following combination of characters: Ventrals 212; vertebrals 217 (taken from parietals to last body scale row that contacts the the most posterior ventral); total vertebrals (body plus tail) 273; subcaudals 56 divided; anal 2; mid-body scale rows 17; body scale rows within 10 vertebrals of anal, 15; neck scale rows 19 at 10th vertebral, 17 at 10th ventral; temporal row 11; first supralabial row 13; second supralabial row 13; third supralabial row 17; supralabials 6-6, 3rd and 4th suborbital; infralabials 6-6. Measurements of Holotype (mm): Snout to vent length 820.0; vent to tail 135.0. Colouration of Holotype (in alcohol): Uniform brown on first third of dorsum, followed by a lighter brown with each scale edged with dark brown forming a distinct reticulated pattern. The head is light brown (lightest on the snout) with a darker interocular 'bar'; nape colour same as snout. There is an incomplete 'V' series of 5 jet black scales on the neck.” (Wells & Wellington 1985). 
CommentDiemenia aspidorhyncha MCCOY 189 and Pseudonaja mengdeni WELLS & WELLINGTON 1985 have been removed from the synonymy of P. nuchalis by SKINNER 2009.

EtymologyNamed for Dr Gregory Mengden, of the Australian Museum, Sydney in recognition of his researches on reptilian cytology. 
  • Brennan, Karl E. C., Morley, Terry, Hutchinson, Mark, and Donnellan, Steve 2012. Redescription of the western desert taipan, Oxyuranus temporalis (Serpentes : Elapidae), with notes on its distribution, diet and genetic variation. Australian Journal of Zoology 59: 227–235 - get paper here
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp. - get paper here
  • Dittmer, D. E., Chapman, T. L. & Bidwell, J. R. 2020. In the shadow of an iconic inselberg: Uluru’s shadow influences climates and reptile assemblage structure at its base. J. Arid Environ. 181, 104179 (2020) - get paper here
  • Hallermann, J. 2020. An annotated list of reptiles and amphibians from the 1905 Hamburg expedition to southwest Australia deposited in the Zoological Museum Hamburg. Evolutionary Systematics 4: 61 - get paper here
  • Mirtschin, P., Rasmussen, A.R. & Weinstein, S.A. 2017. Australia’s Dangerous snakes. CSIRO Publishing, 424 pp. - get paper here
  • O’Shea, M. 2018. The Book of Snakes. Ivy Press / Quarto Publishing, London, - get paper here
  • Skinner, Adam 2009. A multivariate morphometric analysis and systematic review of Pseudonaja (Serpentes, Elapidae, Hydrophiinae). Zoological Journal of the Linnean Society 155 (1): 171-197 - get paper here
  • Swan, G.; Sadlier, R.; Shea, G. 2017. A field guide to reptiles of New South Wales. Reed New Holland, 328 pp.
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wells, R. W. and Wellington, C. R. 1985. A classification of the Amphibia and Reptilia of Australia. Australian Journal of Herpetology, Supplementary Series (1): 1-61 [sometimes cited as 1983] - get paper here
  • Wells, Richard W. 2002. Taxonomy of the Genus Pseudonaja (Reptilia: Elapidae) in Australia. Australian Biodiversity Record (7): 1-41
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
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