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Ptyas mucosa (LINNAEUS, 1758)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Dhaman, Oriental Ratsnake
G: Gebänderte Rattenschlange, Asiatische Rattenschlange
Chinese: 滑鼠蛇 
SynonymColuber mucosus LINNAEUS 1758: 226
Natrix mucosa — LAURENTI 1768
Coluber blumenbachii MERREM 1820: 119
Coluber dhumna CANTOR 1839
Ptyas blumenbachii — FITZINGER 1843
Coryphodon blumenbachii — DUMÉRIL & BIBRON 1854: 184
Leptophis trifrenatus HALLOWELL 1861: 503
Ptyas mucosus — COPE 1861
Ptyas mucosus — GÜNTHER 1864: 249
Zamenis mucosus — BOULENGER 1890
Zamenis mucosus — BOULENGER 1893: 385
Ptyas mucosus — STEJNEGER 1907: 345
Zamenis mucosus — WALL 1907
Zaocys mucosus — WALL 1921: 172
Ptyas mucosus — SMITH 1943: 159
Ptyas mucosus nigricans [CHERNOV 1949]
Ptyas mucosus maximus DERANIYAGALA 1955
Ptyas mucosus — TAYLOR 1965: 734
Ptyas mucosus — MANTHEY & GROSSMANN 1997: 386
Coluber mucosus — LAZELL 1998
Ptyas mucosus nigricans — SHAMMAKOV et al. 1993
Ptyas mucosus — COX et al. 1998: 54
Ptyas mucosus — PINOU & DOWLING 2000
Coluber mucosus — LAZELL 2002
Ptyas mucosa — DAVID & DAS 2004
Ptyas mucosus — SHARMA 2004
Ptyas mucosus — ZHAO 2006
Ptyas mucosa — WALLACH et al. 2014: 617
Ptyas mucosa — WANG et al. 2022 
DistributionAfghanistan, Bangladesh, Burma (Myanmar), Cambodia,
China (Chekiang, Hupeh, Jiangxi, Fujian, Guangdong, Hainan, Guangxi, Yunnan, Xizang, Hong Kong, Hunan),
India (Assam, Mizoram, Tamil Nadu, Kerala, Madhya Pradesh, Himachal Pradesh, Jammu and Kashmir, Maharashtra, Telangana, Karnataka (Castle Rock) [A. Captain, pers. comm.], Punjab [Dino Aulakh, pers. comm.], Gujarat, Tripura, Odisha), Andamans, Sri Lanka, Bhutan,
Indonesia (Sumatra, Java, Bali),
Iran, Laos (Bolikhamxai, Champasak, Khammouan, Louangphrabang, Vientiane and Xiangkhoang), West Malaysia, Nepal, Myanmar, Pakistan, Taiwan, Thailand, Turkmenistan, U.S.S.R., Vietnam (Hoa Binh)

maximus: Sri Lanka

nigricans: Turkmenistan

Type locality: India  
Reproductionoviparous 
TypesHolotype: NRM (NHRM), Stockholm
Holotype: USNM 7510 [Leptophis trifrenatus] 
DiagnosisDiagnosis (genus): Large snakes, with head distinct from the more or less cylindrical body; eye large to very large, the pupil round; scales in 17-15 rows (more rarely 18, 16, 14); subeaudals, paired; apical pits present; normally two or three loreals; a presubocular usually present. Maxillary teeth, 20 to 28, increasing slightly in length and thickness posteriorly. (Taylor 1965: 730)]


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CommentType species: Coluber blumenbachii MERREM 1820 is the type species of the genus Ptyas FITZINGER 1843. According to WALLACH et al. 2014: 615, Coluber mucosus LINNAEUS 1758 is the type species of Ptyas.

Distribution: Possibly in Bhutan (Lenz 2012). For a map see Sindaco et al. 2013.

Habitat: partly arboreal (Harrington et al. 2018).

Genome: Wang et al. 2023 
EtymologyNamed after Latin “mucosus” = slimy, based on the popular notion that snakes are slimy.

Etymology (genus): from Greek “ptyas” = spitter. The gender of Ptyas is feminine (David & Das 2004). 
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