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Ptyas nigromarginata (BLYTH, 1854)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Common NamesE: Green Rat Snake, Black-bordered Rat Snake
G: Grüne Gekielte Rattennatter 
SynonymColuber nigromarginatus BLYTH 1854
Zaocys nigromarginatus — GÜNTHER 1864
Zaocys nigromarginatus — BOULENGER 1893: 376
Zaocys nigromarginatus — WALL 1908: 325
Zaocys dhumnades nigromarginatus — ANGEL & BOURRET 1933
Zaocys nigromarginatus — SMITH 1943: 165
Zaocys nigromarginatus — SWAN & LEVITON 1962: 116
Coluber nigromarginatus — DAS 1996: 58
Ptyas nigromarginata — DAVID & DAS 2004
Zaocys nigromarginatus — SHARMA 2004
Ptyas nigromarginata — WHITAKER & CAPTAIN 2004
Ptyas nigromarginatus — PYRON & BURBRINK 2013
Ptyas nigromarginata — WALLACH et al. 2014: 617 
DistributionNepal, India (Darjelling, Sikkim, Assam ?, Meghalaya, Arunachal Pradesh, Nagaland), N Myanmar (= Burma), N Vietnam (Lao Cai Province, Lang Son, Ha Tay, Kon Tum Province, Ha Giang), Thailand (Nan), Bhutan, Bangladesh,
China (Guizhou, Yunnan, SW Sichuan, SE Xizang = Tibet), elevation 500-2290 m.

Type locality: Vicinity of Darjeeling, India  
TypesHolotype: ZSI 7343 
CommentSynonymy: Lazell et al. (1991) and Lazell (1998) formally synonymized Ptyas and Coluber but at least the latter doesn’t explicitely mention Ptyas nigromarginatus.

Similar species: juveniles are strikingly similar to the juveniles of the species P. dhumnades.

Distribution: Reports from Afghanistan are most likely in error (Vogel & Hauser 2013). These authors listed many localities. 
  • Bai, Linzhuang and Kai Wang. 2015. Ptyas nigromarginata (green ratsnake) diet. Herpetological Review 46 (3): 453 - get paper here
  • Bauer A M; Gunther R 1992. A preliminary report on the reptile fauna of the Kingdom of Bhutan with the description of a new species of scincid lizard (Reptilia: Scincidae). Asiatic Herpetological Research 4: 23-36 - get paper here
  • Blyth, EDWARD. 1855. Notices and descriptions of various reptiles, new or little known [part 2]. J. Asiatic Soc. Bengal, Calcutta, 23 (3): 287-302 [1854] - get paper here
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Das, I. 2012. A Naturalist's Guide to the Snakes of South-East Asia: Malaysia, Singapore, Thailand, Myanmar, Borneo, Sumatra, Java and Bali. Oxford J, ohn Beaufoy Publishing - get paper here
  • David, P. & I. Das 2004. On the grammar of the gender of Ptyas Fitzinger, 1843 (Serpentes: Colubridae). Hamadryad 28 (1&2): 113-116
  • Dowling, H.G., & Jenner, J.V. 1988. Snakes of Burma: checklist of reported species and bibliography. Smithsonian Herp. Inf. Serv. (76): 19 pp. - get paper here
  • Kästle , W., Rai, K. & Schleich, H.H. 2013. FIELD GUIDE to Amphibians and Reptiles of Nepal. ARCO-Nepal e.V., 625 pp. - get paper here
  • Lazell,J.D. 1998. Morphology and the status of the snake genus "Ptyas". Herpetological Review 29 (3): 134 - get paper here
  • Lazell,J.D., Keirans,J.E. & Samuelson,G.A. 1991. The Sulawesi black racer, Coluber (Ptyas) dipsas, and a remarkable ectoparasitic aggregation. Pacific Science 45 (4): 355-361
  • Lenz, Norbert 2012. Von Schmetterlingen und Donnerdrachen - Natur und Kultur in Bhutan. Karlsruher Naturhefte 4, Naturkundemuseum Karlsruhe, 124 pp.
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • O’Shea, M. 2018. The Book of Snakes. Ivy Press / Quarto Publishing, London, - get paper here
  • Pyron, R. Alexander; Frank T. Burbrink 2013. Early origin of viviparity and multiple reversions to oviparity in squamate reptiles. Ecology Letters 17 (1): 13–21 (published online 2013, in print 2014), DOI: 10.1111/ele.12168 - get paper here
  • Sharma, R. C. 2004. Handbook Indian Snakes. AKHIL BOOKS, New Delhi, 292 pp.
  • Smith, M.A. 1943. The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp.
  • Swan, L.W., & Leviton, A.E. 1962. The herpetology of Nepal: a history, check list, and zoogeographical analysis of the herpetofauna. Proc. Cal. Acad. Sci. 32 (6) (4.s.): 103-147. - get paper here
  • Tshewang, S. & L. Letro 2018. The herpetofauna of Jigme Singye Wangchuck National Park in central Bhutan: status, distribution and new records. Journal of Threatened Taxa 10(11): 12489–12498; - get paper here
  • VOGEL, Gernot and Sjon HAUSER 2013. Addition of Ptyas nigromarginata (Blyth, 1854) (Squamata: Colubridae) to the Snake Fauna of Thailand with Preliminary Remarks on Its Distribution. Asian Herpetological Research 4 (3): 166–181
  • Wall,F. 1908. Notes on a collection of snakes from the Khasi Hills, Assam. J. Bombay nat. Hist. Soc. 18: 312-337 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Whitaker, Romulus and Ashok Captain 2004. Snakes of India. Draco Books, 500 pp., reprinted 2007 - get paper here
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