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Ptychozoon kabkaebin GRISMER, WOOD, GRISMER, QUAH, THY, PHIMMACHAK, SIVONGXAY, SEATEUN, STUART, SILER, MULCAHY, ANAMZA & BROWN, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesLao Parachute Gecko 
SynonymPtychozoon kabkaebin GRISMER, WOOD, GRISMER, QUAH, THY, PHIMMACHAK, SIVONGXAY, SEATEUN, STUART, SILER, MULCAHY, ANAMZA & BROWN 2019: 175 
DistributionN/C Laos (Bolikhamxay), likely in adjacent areas of central Vietnam

Type locality: Houay Ta Ang Stream, Pakkading District, Bolikhamxay Province, Laos (18.32997°N, 103.99140°E, 163 m above sea level).  
Reproduction 
TypesHolotype. NCSM 80585 adult female collected by Sengvilay Seateun and Misan Keooudone on 16 June 2012. Paratypes. FMNH 271140 adult female collected by Bryan L. Stuart, Somphouthone Phimmachak, and Niane Sivongxay on 28 May 2007 from Nakai-Nam Theun National Protected Area, Phou Ack Mountain, Boualapha District, Khammouan Province, Laos (17.64433°N, 105.73667°E, 980 m above sea level); and NUOL 00036 adult female collected by Bryan L. Stuart, Somphouthone Phimmachak, and Jennifer A. Sheridan on 17 May 2013 from Nakai-Nam Theun National Protected Area, Phou Ack Mountain, Boualapha District, Khammouan Province, Laos (17.64256 °N, 105.73608 °E, 992 m above sea level). 
DiagnosisDiagnosis. Ptychozoon kabkaebin sp. nov. differs from all other species of Ptychozoon by having the following unique combination of characters: a maximum SVL of 95.4 mm; supranasals not in contact; 8–10 supralabials; 10–12 infralabials; infra-auricular cutaneous flap; weak, supra-auricular ridge present; no dorsal or caudal tubercles; imbricate parachute support scales on dorsal surface of patagia; no prominently raised ridges on ventral surface of patagia; 78–82 midbody dorsal scales; 33–38 ventral scales; an emargination between the pre-antebrachial flap and digit I; no enlarged femoral scales; 22 or 23 enlarged precloacal scales; 5–7 rows of enlarged post-precloacal scales; 13–17 transverse subdigital lamellae on fourth toe; no enlarged dorsal caudal scales forming whorls; approximately 27–30 scales across widest portion of caudal flap; distal lobes fusing to form a short, narrow caudal flap; edges of caudal flap weakly crenulated; caudal lobes angled posteriorly; caudal lobes decrease posteriorly in size; thick, dark, postorbital stripe; four dark body bands between limb insertions; irregularly shaped, white, vertebral markings variably present; and subcaudal region banded in adults (Tables 4, 5).

Comparisons: Differences between Ptychozoon kabkaebin sp. nov. and P. cicakterbang sp. nov. are listed above in the comparisons section of P. cicakterbang sp. nov. Ptychozoon kabkaebin sp. nov. differs from P. intermedium, P. kuhli, and P. trinotaterra in lacking, as opposed to having, caudal tubercles. From P. intermedium, P. nicobarense, P. rhacophorus, P. trinotaterra, and P. kaengkrachanense it differs in having four body bands as opposed to 0–3. Ptychozoon kabkaebin sp. nov. differs from P. bannaense, P. horsfieldii, P. intermedium, P. kuhli, P. nicobarense, P. rhacophorus, P. trinotaterra, and P. kaengkrachanense in having an emarginated preantebrachial flap as opposed to lacking an emargination. From P. popaense it differs by having a maximum SVL of 95.4 mm versus 86.2 mm. Ptychozoon kabkaebin sp. nov. differs from P. lionotum, and P. tokehos sp. nov. by having a significantly higher mean number of infralabials. Ptychozoon kabkaebin sp. nov. differs from P. tokehos sp. nov. in having a significantly wider head. Even though Ptychozoon kabkaebin sp. nov. and P. tokehos sp. nov. have nearly discrete differences in their numbers of midbody scales (78–82 and 80–95, respectively), their mean differences (80.3 versus 89.0) were not significantly different in the ANOVA (p=0.09) although they were significantly different in a two-sample Student t-test (t=5.585; p=0.002). Ptychozoon kabkaebin sp. nov. is the sister species of P. lionotum but differs further from it by having a significantly shorter snout. Ptychozoon kabkaebin sp. nov. is well-separated from P. cicakterbang sp. nov. and P. lionotum in the PCA and from all species in the DAPC where their 95% confidence ellipses do not overlap. Ptychozoon kabkaebin sp. nov. occupies a significantly different position along PC1 from that of P. cicakterbang sp. nov. and along PC2, it occupies a significantly different positon from that of P. tokehos sp. nov. From all species of the lionotum group it differs by having an uncorrected pairwise sequence divergence of 4.1–14.4%. Combinations of other characters differentiating P. kabkaebin sp. nov. from the other more distantly related species are presented in Tables 4, 5; Figs. 3, 5, 6 in Grismer et al. 2019. 
CommentHabitat: large boulders in a mosaic of evergreen, deciduous, and pine forest. NCSM 80585 was collected during the day from beneath a house on stilts that was located along a stream in degraded semi-evergreen forest at 160 m in elevation. FMNH 271140 was collected during the day (1115 h) at 980 m in elevation and NUOL 00036 was collected during the night (2130 h) at 992 m in elevation, both on large boulders in a mosaic of evergreen, deciduous, and pine forest with grassy understory at the edge of a high cliff on a steep escarpment. 
EtymologyThe specific epithet kabkaebin is the Lao word used for Ptychozoon. 
References
  • GRISMER, L. LEE; PERRY L. WOOD JR., JESSE L. GRISMER, EVAN S. H. QUAH, NEANG THY, SOMPHOUTHONE PHIMMACHAK, NIANE SIVONGXAY, SENGVILAY SEATEUN, BRYAN L. STUART, CAMERON B. SILER, DANIEL G. MULCAHY, TASHITSO ANAMZA & RAFE M. BROWN 2019. Geographic structure of genetic variation in the Parachute Gecko Ptychozoon lionotum Annandale, 1905 across Indochina and Sundaland with descriptions of three new species. Zootaxa 4638 (2): 151–198 - get paper here
 
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