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Ptychozoon tokehos GRISMER, WOOD, GRISMER, QUAH, THY, PHIMMACHAK, SIVONGXAY, SEATEUN, STUART, SILER, MULCAHY, ANAMZA & BROWN, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesCambodian Parachute Gecko 
SynonymPtychozoon tokehos GRISMER, WOOD, GRISMER, QUAH, THY, PHIMMACHAK, SIVONGXAY, SEATEUN, STUART, SILER, MULCAHY, ANAMZA & BROWN 2019: 179 
DistributionCambodia (Kampong Speu), Thailand, S Vietnam

Type locality: Kirirom National Park, Phnom Sruoch District, Kampong Speu Province, Cambodia (11.32611°N, 104.06556°E, 700 m in elevation).  
Reproduction 
TypesHolotype. FMNH 261853 adult female, collected by Bryan L. Stuart and Dara Anon 5 June 2000. Paratypes. FMNH 261851–52 bear the same collection data as the holotype except they were collected on 3 June 2000. FMNH 261854 bears the same collection data as the holotype except it was collected on 5 November 2000 by Joe Walston. CBC 03162 adult female collected by Neang Thy on 9 June 2017 from Bokor National Park, Teuk Chou District, Kampot Province, Cambodia (10.60156°N, 104.03200°E, 325 m in elevation). NCSM 98986 adult female collected by Neang Thy on 10 June 2017 from Bokor National Park, Teuk Chou District, Kampot Prov- ince, Cambodia (10.60372°N, 104.02944°E, 328 m in elevation). NCSM 98987 adult female collected by Neang Thy on 12 June 2017 from Bokor National Park, Teuk Chou District, Kampot Province, Cambodia (10.58753°N, 104.03269°E, 315 m in elevation). FMNH 177359 adult male collected by Edward H. Taylor on 11 June 1969 from the Khao Chong Reserve, Trang Province, Thailand (7.54350°N 99.79800°E, 127 m in elevation). FMNH 181844 adult male collected by W. Ronald Heyer on 11 June 1969 from Pak Thong Chai District, Nakhon Ratchasima Prov- ince, Sakaerat (Environmental Research Station), Thailand (14.50000°N, 100.86670°E, 16 m in elevation). 
DiagnosisDiagnosis. Ptychozoon tokehos sp. nov. differs from all other species of Ptychozoon by having the following unique combination of characters: a maximum SVL of 97.5 mm; supranasals not in contact; 8–11 supralabials; 10–12 infralabials; infra-auricular cutaneous flap; weak supra-auricular ridge present; no dorsal or caudal tubercles; imbricate parachute support scales on dorsal surface of patagia; prominently raised ridges on ventral surface of patagia; 80–95 midbody dorsal scales; 30–37 ventral scales; an emargination between the pre-antebrachial flap and digit I; no enlarged femoral scales; 20–24 pore-bearing precloacal scales in males; 18–25 enlarged precloacal scales; 4–7 rows of enlarged post-precloacal scales; 13–18 transverse subdigital lamellae on the fourth toe; approximately 28–34 scales across the widest portion of the caudal flap; enlarged dorsal caudal scales forming intermittent whorls; distal lobes fusing to form a short, narrow, caudal flap; edges of caudal flap smooth; caudal lobes angled posteri- orly; caudal lobes variably showing posterior reduction in size; postorbital striping variable; four dark body bands between limb insertions; and irregularly shaped, white, vertebral markings usually not present (absent in 14 of 16 specimens; Tables 4, 5).

Comparisons. Differences between Ptychozoon tokehos sp. nov., P. kabkaebin sp. nov., and P. cicakterbang sp. nov. are listed above in the comparisons sections of those species. Ptychozoon tokehos sp. nov. differs from P. intermedium, P. kuhli, and P. trinotaterra in lacking, as opposed to having, caudal tubercles. From P. intermedium, P. nicobarense, P. rhacophorus, P. trinotaterra, and P. kaengkrachanense it differs in having four body bands as opposed to 0–3. Ptychozoon tokehos sp. nov. differs from P. bannaense, P. horsfieldii, P. intermedium, P. kuhli, P. nicobarense, P. rhacophorus, P. trinotaterra, and P. kaengkrachanense in having an emargination between the pre-antebrachial flap and digit I as opposed to no emargination. From P. popaense it dif- fers by having a maximum SVL of 97.5 mm versus 86.2 mm. Ptychozoon tokehos sp. nov. is well-separated from P. cicakterbang sp. nov. and P. lionotum in the PCA and from all species in the DAPC where their 95% confidence ellipses do not overlap. Ptychozoon tokehos sp. nov. occupies a significantly different position along PC1 from that of P. cicakterbang sp. nov. and along PC2, it occupies a significantly different positon from those of P. kabkaebin sp. nov. and P. lionotum. From all other species of the P. lionotum group it is further separated by an uncorrected pairwise sequence divergence of 4.1–15.5%. Combinations of other characters differentiating P. kabkaebin sp. nov. from other more distantly related species are presented in Tables 4, 5; Figs. 3, 5, 6 in Grismer et al. 2019. 
CommentHabitaT: forest in hilly areas from sea level to at least 700 m in elevation. This species does quite well in disturbed forests and is commonly found on man-made structures. The holotype and paratypes from Kirirom National Park (FMNH 261851–54) were all found during the day (1400–1630 h) on or near the exterior walls of a building on a grassy plateau within open pine forest. FMNH 261851 was found on the branch of a large tree abutting an exterior wall, FMNH 261852 was found beneath a layer. of paint peeling off the exterior wall of the building approximately 2.5 m above the ground, and FMNH 261853–54 were found on the exterior walls. 
EtymologyThe specific epithet tokehos is the Khmer (Cambodian) word used for Ptychozoon. 
References
  • GRISMER, L. LEE; PERRY L. WOOD JR., JESSE L. GRISMER, EVAN S. H. QUAH, NEANG THY, SOMPHOUTHONE PHIMMACHAK, NIANE SIVONGXAY, SENGVILAY SEATEUN, BRYAN L. STUART, CAMERON B. SILER, DANIEL G. MULCAHY, TASHITSO ANAMZA & RAFE M. BROWN 2019. Geographic structure of genetic variation in the Parachute Gecko Ptychozoon lionotum Annandale, 1905 across Indochina and Sundaland with descriptions of three new species. Zootaxa 4638 (2): 151–198 - get paper here
 
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