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Ptyodactylus puiseuxi BOUTAN, 1893

IUCN Red List - Ptyodactylus puiseuxi - Least Concern, LC
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Higher TaxaPhyllodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Levant fan-footed gecko, Israeli Fan-fingered Gecko 
SynonymPtyodactylus puiseuxi BOUTAN 1893: 379
Ptyodactylus lobatus syriacus PERACCA 1894
Ptyodactylus lobatus sancti-montis BARBOUR 1914
Ptyodactylus hasselquistii puiseuxi — WERNER 1965
Ptyodactylus hasselquistii puisieuxi [sic] — ESTERBAUER 1985
Ptyodactylus puiseuxi — KLUGE 1993
Ptyodactylus puiseuxi — RÖSLER 2000: 107
Ptyodactylus puiseuxi — MEIRI et al. 2019
Ptyodactylus puiseuxi — TAMAR et al. 2025 
DistributionIsrael, Syria, Jordan (Palmyra Azraq), Iraq, Lebanon

Type locality: Lac de Houleh, Syria. Neotype locality: from Gonen, Hula Valley, Israel (33.12297°N, 35.64412°E; ca. 100m a.s.l.).

sancti-montis: Type locality: Rasheya, base of Mt. Hermon, Syria

syriacus: Type locality: Dscherasch, Est-Giordano = Jerash, Jordan  
Reproductionoviparous 
TypesNeotype: SMNHTAU:R. 19711, adult male, collected on the 30 June 2020 by G. Stark, R. Schwarz, and K. Tamar; original type was MHNB, but now lost (Werner & Sivan 1994, DE LISLE et al. 2013: 221, Tamar et al. 2025)
Holotype: MCZ 9757 [sancti-montis]
Holotype: MSNTO R238 [syriacus]
Holotype: lost, fide Tamar et al. 2025 [barroisi]
Holotype: lost, fide Tamar et al. 2025 [bischoffsheimi]
Holotype: lost, fide Tamar et al. 2025 [montmahoui] 
DiagnosisDiagnosis: A middle-sized Ptyodactylus species with a large triangular head, tail shorter than body length; dorsal scales small, granular and mixed with smooth, round and small conical tubercles. No tubercles in front of the head. Tail shorter than snout-vent length, with 4 dark bands and small white spots. No caudal whorl. Snout-vent length 63 mm, tail length 54 mm. Colour of specimens from Ana preserved in formaldehyde: light brown with white spots. The Rutba material is darker and more robust.


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CommentSynonymy after KLUGE 1993 (who used both BOUTAN and BOUTON several times although his bibliography lists only papers by BOUTAN), and Tamar et al. 2025. Color image in IN DEN BOSCH (1999). P. puiseuxi and its synonyms have been listed as synonyms of P. hasselquistii by WERMUTH (1965).

Distribution: see Bar et al. 2021, METALLINOU et al. 2015: 210 (Fig. 1), and Tamar et al. 2025 for maps.

Reference images: see Uetz et al. 2024 for high-resolution reference images for this species. 
EtymologyNamed after Victor Alexandre Puiseux (1820-1883), an astronomer. On graduating (1837) he entered Ecole Normale Superieure, where he was awarded his doctorate (1841). 
References
  • Afrasiab, Saman R. and Sarbaz I. Mohamad 2009. A study on cave-dwelling geckos in Iraq, with the description of a new species from Saffine mountain (Reptilia: Gekkonidae). Zoology in the Middle East 47:49-56 - get paper here
  • Al-Quran, S. 2009. The Herpetofauna of the Southern Jordan. American-Eurasian J. Agric. & Environ. Sci., 6 (4): 385-391 [this journal has a dubious record, see http://goo.gl/81SBPv]
  • Aloufi, A., Amr, Z., & Baker, M. A. 2022. Reptiles from'Uruq Bani Ma'arid and Harat al Harrah protected areas in Saudi Arabia: Reptiles from two protected areas in Saudi Arabia. Herpetology Notes 15: 483-491 - get paper here
  • Bar, A., Haimovitch, G. and Meiri, S. 2021. Field guide to the amphibians and reptiles of Israel. Edition Chimaira, Frankfurt Am Main. ISBN 9783-89973-120-0 - get paper here
  • Bar, Aviad and Guy Haimovitch 2012. A Field Guide to Reptiles and Amphibians of Israel. Pazbar LTD, 246 pp. - get paper here
  • Barbour, Thomas 1914. Notes on some reptiles from Sinai and Syria. Proc. New England zool. Club 5: 73-92 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • BOSCH, H.A.J. In DEN 1999. Die Eidechsen des Mount Hermon im Biotop mit ersten Daten zur Fortpflanzung. Elaphe 7 (4): 68-75
  • Boutan, L. 1893. Mémoire sur les reptiles rapportés de Syrie par le Docteur Théod. Barrois. 1re Partie. Genre Ptyodactyle. Revue Biologique du Nord de la France, 5:1-36 [reprint pagination]. - get paper here
  • Disi, A.M.; Modry, D.; Nečas, P. & Rifai, L. 2001. Amphibians and reptiles of the Hashemite Kingdom of Jordan. Edition Chimaira, Frankfurt, 408 pp.
  • Disi, Ahmad M., Zuhair S. Amr and Harald Martens 2004. On a collection of amphibians and reptiles made by J. Klapperich in Jordan. Herpetozoa 16 (3/4):141-150 - get paper here
  • Esterbauer, H. 1985. Zur Herpetofauna Südwestsyriens. Herpetofauna 7 (38): 23-34 - get paper here
  • Goldberg, S.R. & Maza, E. 2017. Ptyodactylus puiseuxi (Israeli Fan-fingered Gecko) Reproduction. Herpetological Review 48 (4): 853-854 - get paper here
  • Heimes, P. 1987. Beitrag zur Systematik der Fächerfinger (Sauria: Gekkonidae: Ptyodactylus). Salamandra 23 (4): 212-235 - get paper here
  • Hraoui-Bloquet, S.; Sadek, R.A.; Sindaco, R. & Venchi, A. 2002. The herpetofauna of Lebanon: new data on distribution. Zoology in the Middle East 27: 35-46 - get paper here
  • Leviton,A.E.; Anderson,S.C.; Adler, K.; Minton,S.A. 1992. Handbook to Middle East Amphibians and Reptiles. SSAR, Oxford, Ohio (Contr. to Herpetol. No. 8), 1-252
  • Lisle, H.F. de; Nazarov, R.A.; Raw, L.R.G. & Grathwohl, J. 2013. Gekkota: a catalogue of recent species. [type catalogue] Privately published, 387 pp.
  • Meiri, Shai; Amos Belmaker, Daniel Berkowic, Kesem Kazes, Erez Maza, Guy Bar-Oz and Roi Dor 2019. A checklist of Israeli land vertebrates. Israel Journal of Ecology and Evolution - get paper here
  • Metallinou, Margarita; Jan Cervenka, Pierre-Andre Crochet, Lukas Kratochvil, Thomas Wilms, Philippe Geniez, Mohammed Y. Shobrak, Jose C. Brito, Salvador Carranza 2015. Species on the rocks: Systematics and biogeography of the rock-dwelling Ptyodactylus geckos (Squamata: Phyllodactylidae) in North Africa and Arabia. Molecular Phylogenetics and Evolution 85: 208-220 - get paper here
  • Modrý, P., Rifai, L., Abu Baker, M. & Amr, Z. 2004. Amphibians and reptiles of the Hashemite Kingdom of Jordan. Denisia 14 (Neue Serie 2): 407-420 - get paper here
  • Peracca, M. G. 1894. Viaggio del Dr. E. Festa in Palestina, nel Libano e regioni vicine. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino, 9: 1-20 - get paper here
  • Pola, L., Hejduk, V., Winkelhöfer, T., Zíka, A., Baker, M. A. A., & Amr, Z. S. 2020. Recent Observations on Amphibians and Reptiles in the Hashemite Kingdom of Jordan. Jordan Journal of Natural History, 7: 11-29 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Sindaco R., Fedrighini, N. & VEnchi, A. 1995. Contribution to the herpetology of Jordan. Boll. Mus. Reg. Sci. Nat. Torino 13 (1): 389-405
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Sindaco, R.; Serra, G. & Menegon, M. 2006. New data on the Syrian herpetofauna with a newly-recorded species of snake. Zoology of the Middle-East 37: 29-38 - get paper here
  • Tamar, K., Carranza, S., Šmíd, J., & Moravec, J. 2025. Taxonomy and phylogeny of the Levant fan-footed gecko Ptyodactylus puiseuxi Boutan, 1893 and the resurrection of Ptyodactylus bischoffsheimi Boutan, 1893 (Squamata, Phyllodactylidae). Systematics and Biodiversity, 23(1), 2454011 - get paper here
  • Uetz, P.H.; Patel, M.; Gbadamosi, Z.; Nguyen, A.; Shoope, S. 2024. A Reference Database of Reptile Images. Taxonomy 4: 723–732 - get paper here
  • Werner Y L. Sivan N. 1996. Systematics and zoogeography of Ptyodactylus (Reptilia: Sauria: Gekkonidae) in the levant: 3, experimental and natural hybrids of P. guttatus and P. puiseuxi. Israel Journal of Zoology. 1996. vol. 42, no. 2, pp. 185-202. - get paper here
  • Werner, Y. L. 1965. Über die israelischen Geckos der Gattung Ptyodactylus und ihre Biologie. Salamandra 1 (1-2): 15-25 - get paper here
  • Werner, Yehudah L.;Sivan, Naomi 1993. Systematics and zoogeography of Ptyodactylus (Reptilia: Sauria: Gekkonidae) in the Levant:1. Biometry of three species in Israel. Revista Española de Herpetología 7: 47-64
  • Werner, Yehudah L.;Sivan, Naomi 1994. Systematics and zoogeography of Ptyodactylus (Reptilia: Sauria: Gekkonidae) in the levant : 2, taxonomy, with a review of ecology and zoogeography. Revista Española de Herpetologia 8: 105-122
 
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