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Rena humilis BAIRD & GIRARD, 1853

IUCN Red List - Rena humilis - Least Concern, LC

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Higher TaxaLeptotyphlopidae, Epictinae, Epictini, Typhlopoidea, Serpentes (snakes) 
SubspeciesRena humilis boettgeri (WERNER 1899)
Rena humilis cahuilae KLAUBER 1939
Rena humilis chihuahuaensis TANNER 1985
Rena humilis dugesii (BOCOURT 1881)
Rena humilis humilis (BAIRD & GIRARD 1853)
Rena humilis levitoni MURPHY 1975 (fide LINER 1994)
Rena humilis lindsayi MURPHY 1975 (fide LINER 1994)
Rena humilis segregus KLAUBER 1939
Rena humilis tenuiculus (GARMAN 1883)
Rena humilis utahensis TANNER 1938 
Common NamesE: Western Threadsnake
cahuilae: Desert Blind Snake, Desert Threadsnake
humilis: Southwestern Blind Snake/Threadsnake
segregus: Trans-Pecos Blind Snake/Threadsnake
utahensis: Utah Blind Snake/Threadsnake
G: Mexikanische Schlankblindschlange 
SynonymRena humilis BAIRD & GIRARD 1853: 143
Stenostoma humile - PETERS 1857
Catadon dugesii BOCOURT 1881
Catodon dugesii BOCOURT 1881
Siagonodon dugesii - BOCOURT 1882: 507
Stenostoma tenuiculum GARMAN 1884: 5
Rena dugesii - COPE 1887: 64
Rena tenuicula - COPE 1887: 91
Glauconia humilis - BOULENGER 1893: 70
Siagonodon humilis - VAN DENBURGH 1897
Glauconia boettgeri WERNER 1899: 116
Leptotyphlops humilis - RUTHVEN 1907
Leptotyphlops humilis humilis - KLAUBER 1931
Leptotyphlops humilis slevini KLAUBER 1931
Leptotyphlops dugesii - TAYLOR 1940: 143
Leptotyphlops humilis dugesi - KLAUBER 1940
Leptotyphlops humilis tenuiculus - KLAUBER 1940: 538
Leptotyphlops chumilis RHODES (error typographicus) 1966
Leptotyphlops humilis boettgeri - SMITH & LARSEN 1974: 93
Leptotyphlops humilis levitoni MURPHY 1975
Leptotyphlops humilis lindsayi MURPHY 1975
Leptotyphlops humilis — STEBBINS 1985: 171
Leptotyphlops humilis — MCDIARMID, CAMPBELL & TOURÉ 1999: 32
Rena humilis — ADALSTEINSSON, BRANCH, TRAPE, VITT & HEDGES 2009
Rena humilis — CROTHER et al. 2012

Leptotyphlops humilis cahuilae KLAUBER 1939
Leptotyphlops humilis cahuilae KLAUBER 1931 (fide CROTHER 2000)
Leptotyphlops humilis cahuilae — CROTHER 2000: 65
Rena humilis cahuilae — CROTHER et al. 2012

Leptotyphlops humilis dugesii (BOCOURT 1881)
Catodon dugesii BOCOURT 1881: 81
Leptotyphlops humilis dugesi — ZWEIFEL 1959
Leptotyphlops humilis dugesi — WEBB 1984
Leptotyphlops humilis dugesii — PETERSON 1995

Leptotyphlops humilis humilis (BAIRD & GIRARD 1853)
Rena humilis BAIRD & GIRARD 1853: 143
Leptotyphlops humilis humilis — CROTHER 2000: 65

Leptotyphlops humilis segregus KLAUBER 1939
Leptotyphlops humilis segregus KLAUBER 1939
Leptotyphlops humilis segregus — CONANT & COLLINS 1991: 145
Leptotyphlops humilis segregus — CROTHER 2000: 65
Leptotyphlops humilis segregus — TENNANT & BARTLETT 2000: 34
Rena humilis segregus — CROTHER et al. 2012

Leptotyphlops humilis utahensis TANNER 1938
Leptotyphlops humilis utahensis TANNER 1938
Leptotyphlops humilis utahensis — CROTHER 2000: 65
Rena humilis utahensis — CROTHER et al. 2012 
DistributionUSA (S California, S Nevada, S Arizona, S New Mexico, S Texas), f
Mexico (Baja California, Sonora, Chihuahua, Sinaloa, N Coahuila, N Durango)

boettgeri: Mexico (Baja California Sur and Cerralvo Island); Type locality: unknown (fide HAHN 1980).

cahuilae: SE California, SW Arizona; Mexico (Baja California); Yaqui Well, San Diego County, California, USA.

chihuahuaensis: Mexico (Chihuahua)

dugesii: Mexico (Sonora, Sinaloa, Jalisco, Colima); Type locality: Colima, Mexico.

humilis: USA (S California, S Nevada, Arizona); Mexico (N Baja California); Type locality: "Valliecitas", California, USA.

levitoni: Mexico (Baja California: Isla Santa Catalina); Type locality: Isla Santa Catalina, Gulf of California, Mexico (26° 40' N., 110° 47' W.).

lindsayi: Mexico (Baja California: Isla Carmen); Type locality: Isla Carmen, Gulf of California, Mexico (25° 57' N., 111° 12' W.).

segregus: USA (SE Arizona, Texas (Trans-Pecos), New Mexico), Mexico (N Coahuila); Type locality: Chalk Draw, Brewster County, Tex.

slevini: Baja California Cape region; Type locality: La Paz, Baja California.

tenuiculus: Mexico (San Luis Potosi, Tamaulipas); Type locality: San Luis Potosi (city?).

utahensis: SW Utah, SE Nevada. Type locality: Saint George, Washington County, Utah, USA. Map legend:
Type locality - Type locality.
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Typescahuilae: Holotype: SDSNH 2637
dugesii: MHNH Paris
humilis: USNM 2101
levitoni: CAS 135146
lindsayi: Holotype: SDSNH 44386.
segregus: USNM 103670
slevini: CAS 53721
tenuiculus: MCZ 4519 
CommentSubspecific classification according to HAHN 1980 and LINER 1994. Leptotyphlops humilis tenuiculus is a synonym of Leptotyphlops dulcis iversoni fide Smith et al. 1998. Grismer (1999) synonymized Leptotyphlops humilis levitoni with Leptotyphlops humilis.

Type species: Rena humilis BAIRD & GIRARD 1853 is the type species of the genus Rena BAIRD & GIRARD 1853.

Synonymy: Kaiser et al. 2013 considered the generic name Evanwhittonus Hoser 2012 invalid and rejected its use instead of Rena.

Diagnosis (genus): Species of Rena have 14 midbody scale rows, 10 (12 rarely) midtail scale rows, 168–312 middorsal scale rows, 9–21 subcaudals, 2–3 supralabials, moderate or large (rarely small) anterior supralabials, 205–389 mm maximum adult total length, a body shape of 26–60 (total length/width), a relative tail length of 3.1–8.6 %, a tail shape of 1.9–3.8, no striped pattern, brown or purplish brown dorsal color, and white venter (Table 2 in ADALSTEINSSON et al. 2009). They also have a small supraocular scale. They are distinguished from the other genus in this subtribe, Tricheilostoma, by having a white (not brown or pale brown) venter, usually two supralabials (three in R. bressoni, R. dissecta, and R. myopica), and in having a higher number (on average) of middorsal scales (Table 2). The support for this group was 100% BP and 100% PP for the four-gene tree (Fig. 3); only one species was included in the nine-gene tree (Fig. 4). (from ADALSTEINSSON et al. 2009). 
EtymologyEtymology (genus): The generic name is feminine and derived from the Latin noun ren (kidney), apparently in allusion to the kidney color (reddish brown) of the type species. 
References
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