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Riama raneyi (KIZIRIAN, 1996)

IUCN Red List - Riama raneyi - Vulnerable, VU

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Higher TaxaGymnophthalmidae (Cercosaurinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Subspecies 
Common Names 
SynonymProctoporus raneyi KIZIRIAN 1996: 126
Proctoporus raneyi — DOAN & SCHARGEL 2003
Riama raneyi — DOAN & CASTOE 2005 
DistributionEcuador, elevation 2000–2910 m

Type locality: 3.3 km (by road) ESE Cuyujua, Napo. Ecuador [78° 00 W, 00° 25’ S], 2350 m elevation.  
Reproductionoviparous 
TypesHolotype: KU 142905, adult male 
DiagnosisDiagnosis:-(1) Frontonasal distinctly longer than to equal to frontal; (2) nasoloreal suture usually absent; (3) supraoculars two or three, usually three, usually second and third in contact with ciliaries; (4) superciliary series incomplete, usually one; (5) supralabial-subocular fusion usually absent; (6) postoculars two or three, usually three; (7) postparietals usually two or three; (8) supratympanic temporals two or three, usually three; (9) genials two or three, usually two, transverse sutures not perpendicular with respect to midline of body; (10) dorsals rectangular, juxtaposed, striated/keeled; (11) longitudinal dorsal scale rows 22-28; (12) transverse dorsal scale rows in males 38-42, in females 33-42; (13) transverse ventral scale rows 19-24; (14) lateral scale rows 2-4, usually two; (15) femoral pores in males 7-10, in females zero or one; scales between femoral pores in males 2-4, in females 16-18; (16) subdigital scales on Toe I 4-6; (17) limbs not overlapping when adpressed against body in adults; (18) anterior cloacal plate scales paired; (19) hemipenis capitate; flounces forming two chevrons; asulcate flounces usually in contact, distinctly narrower than sulcate flounces; asulcate expansion pleat absent; (20) dorsum unicolored dark brown or distinctly bicolored; venter unicolored dark brown or with white along scale many (especially longitudinal) sutures on anterior belly scales.
Proctoporus raneyi can be distinguished by the following (condition for P. raneyi in parentheses): Ecuadorian congeners: P. balneator, P. colomaromani, P. hypostictus, P. labionis, P. meleagris, P. oculatus, P. orcesi, and P. stigmatoral: supraoculars four (two or three). P. anatoloros, P. balneator, P. oculatus, P. orcesi, P. simoterus, P. stigmatoral, P. unicolor, P. vespertinus, and P. vietus: superciliaries two or more (usually one). P. cashcaensis: supratympanic temporals usually two (usually three); supraoculars usually three (usually four); hemipenial flounces in two columns (two chevrons). P. petrorum: frontonasal slightly shorter than (slightly longer than or equal in length to) frontal; femoral pores in males four (7-10); scales between femoral pores eight (2-4). P. luctuosus group: lateral scales in wide (narrow) band, superciliary series complete (incomplete); femoral pores per leg in females two or more (zero or one); dorsolateral stripes and ocelli present (absent). P. achlyens, P. laevis, and P. luctuosus: nasoloreal suture complete (usually absent). P. columbianus: Dorsal scales smooth (striated/keeled), nasoloreal suture complete (usually absent). P. striatus: hemipenis acapitate (capitate); asulcate expansion pleat on hemipenis present (absent). (Kizirian 1996)


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Comment 
EtymologyThe specific epithet is a noun in the genitive singular case formed from the modern name Raney, and is a patronym honoring Richard H. Raney of Lawrence, Kansas, in recognition of his generous support of the Panorama Society, Museum of Natural History, The University of Kansas. Panorama funds were used to support, in part, the research that led to the discovery of this new species. 
References
  • Arteaga, A.; Bustamante, L.; Vieira, J. 2024. Reptiles of Ecuador. Khamai Foundation & Tropical Herping, 1073 pp. - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Doan, T. M. & Castoe, T.A. 2005. Phylogenetic taxonomy of the Cercosaurini (Squamata: Gymnophthalmidae), with new genera for species of Neusticurus and Proctoporus. Zoological Journal of the Linnean Society, 143: 405–416. - get paper here
  • Doan, T.M. 2003. A south-to-north biogeographic hypothesis for Andean speciation: evidence from the lizard genus Proctoporus (Reptilia, Gymnophthalmidae). Journal of Biogeography 30: 361–374 - get paper here
  • Doan, T.M. & Schargel, W.E. 2003. Bridging the gap in Proctoporus distribution: a new species (Squamata: Gymnophthalmidae) from the Andes of Venezuela. Herpetologica 59 (1): 68-75 - get paper here
  • Kizirian, D. A. 1996. A review of Ecuadorian Proctoporus (Squamata: Gymnophthalmidae) with descriptions of nine new species. Herpetological Monographs 10: 85-155 - get paper here
  • Sánchez-Pacheco, Santiago J.; David A. Kizirian, and Pedro M. Sales-Nunes 2011. A New Species of Riama from Ecuador Previously Referred to as Riama hyposticta (Boulenger, 1902) (Squamata: Gymnophthalmidae). American Museum Novitates (3719): 1-15 - get paper here
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
 
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