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Riama orcesi (KIZIRIAN, 1995)

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Higher TaxaGymnophthalmidae (Cercosaurinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Subspecies 
Common Names 
SynonymProctoporus orcesi KIZIRIAN 1995: 66
Proctoporus orcesi — KIZIRIAN 1996: 122
Proctoporus orcesi — DOAN & SCHARGEL 2003
Riama orcesi — DOAN & CASTOE 2005 
DistributionNE Ecuador, elevation 1100–2460 m

Type locality: 3.3 km (by road) ESE Cuyujua, Napo. Ecuador [78° 00 W, 00° 25’ S], 2350 m elevation.  
Reproductionoviparous 
TypesHolotype: KU 146709 
DiagnosisDiagnosis:-(1) Frontonasal distinctly longer to distinctly shorter than frontal; (2) nasoloreal suture usually absent; (3) supraoculars four, usually none in contact with ciliaries; (4) superciliary series usually complete, 4-6; (5) supralabial-subocular fusion absent; (6) postoculars 2-4, usually three; (7) postparietals two or three, usually three; (8) supratympanic temporals 2-4, usually two or three; (9) genials two or three, usually three, transverse sutures nearly perpendicular with respect to midline of body; (10) dorsals rectangular, juxtaposed, with low rounded keel; (11) longitudinal dorsal scale rows 13-23; (12) transverse dorsal scale rows in males 32-43, in females 33-42; (13) transverse ventral scale rows 19-23; (14) lateral scale rows 0-8, usually 4-8; (15) femoral pores in males 10-14, absent in females; scales between femoral pores in males 0-2; (16) subdigital scales on Toe I 4-6; (17) limbs not overlapping when adpressed against body in adults; (18) anterior cloacal plate scales paired; (19) hemipenis small, capitate; flounces bearing spines, forming two chevrons; distal one-third of sulcus spermaticus forked; (20) dorsum brown with distinct pale dorsolateral stripe above forelimbs and hind limbs.
Proctoporus orcesi is unique among Proctoporus in that the hemipenis is relatively small and the distal one-third of the sulcus spermaticus is divided. It can be further distinguished from its congeners by the following characters (condition for P. orcesi in parentheses): P. anatoloros (usually), P. balneator, P. colomaromani, P. columbianus, P. hypostictus, P. labionis, P. luctuosus group, P. meleagris, P. oculatus, P. simoterus, P. unicolor, and P. ventrimaculatus: femoral pores in females present (absent). P. cashcaensis, P. petrorum, P. raneyi, P. stigmatoral, P. unicolor, and P. vespertinus: superciliary series incomplete (complete). P. vietus: dorsal scales rugose (striated/keeled). P. columbianus, P. meleagris, P. simoterus, and P. ventrimaculatus: dorsal scales smooth (striated/keeled).


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Comment 
EtymologyThe species is named in honor of Gustavo Orces V., a pioneer in Ecuadorian natural history. 
References
  • Arteaga, A.; Bustamante, L.; Vieira, J. 2024. Reptiles of Ecuador. Khamai Foundation & Tropical Herping, 1073 pp. - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Doan, T. M. & Castoe, T.A. 2005. Phylogenetic taxonomy of the Cercosaurini (Squamata: Gymnophthalmidae), with new genera for species of Neusticurus and Proctoporus. Zoological Journal of the Linnean Society, 143: 405–416. - get paper here
  • Doan, T.M. 2003. A south-to-north biogeographic hypothesis for Andean speciation: evidence from the lizard genus Proctoporus (Reptilia, Gymnophthalmidae). Journal of Biogeography 30: 361–374 - get paper here
  • Doan, T.M. & Schargel, W.E. 2003. Bridging the gap in Proctoporus distribution: a new species (Squamata: Gymnophthalmidae) from the Andes of Venezuela. Herpetologica 59 (1): 68-75 - get paper here
  • Kizirian, D. A. 1996. A review of Ecuadorian Proctoporus (Squamata: Gymnophthalmidae) with descriptions of nine new species. Herpetological Monographs 10: 85-155 - get paper here
  • Kizirian, D.A. 1995. A new species of Proctoporus (Squamata: Gymnophthalmidae) from the Andean Cordillera Oriental of northeastern Ecuador. Journal of Herpetology 29 (1): 66-72 - get paper here
  • Sánchez-Pacheco, Santiago J.; David A. Kizirian, and Pedro M. Sales-Nunes 2011. A New Species of Riama from Ecuador Previously Referred to as Riama hyposticta (Boulenger, 1902) (Squamata: Gymnophthalmidae). American Museum Novitates (3719): 1-15 - get paper here
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
 
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