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Sanzinia volontany VENCES & GLAW, 2004

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Higher TaxaBoidae (Sanziniinae, Sanziniidae), Henophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesE: Nosy Komba ground boa, Madagascar Tree Boa
G: Nosy Komba-Hundskopfboa 
SynonymSanzinia madagascariensis volontany VENCES & GLAW 2004
Sanzinia madagascariensis volontany — D’CRUZE et al. 2010
Sanzinia volontany — REYNOLDS et al. 2014
Sanzinia volontany — HYDE ROBERTS et al. 2014 
DistributionW Madagascar (Nosy Komba)

Type locality: Ampijoroa (Ankarafantsika Reserve)  
Reproduction 
TypesHolotype: ZSM 804/2001, adult male; paratypes: MNHN 1978.96 and 1978.97, subadult 
CommentSubspecies: Both (sub-) species are parapatric and their distribution areas approach each other only in the very south of Madagascar. Some authors suggested that S. m. volontany should be elevated to specific status despite an apparent lack of diagnostic morphological differences (Orozco-Terwengel et al., 2008, Reynolds et al. 2014).

Synonymy: Listed as synonym of Sanzinia madagascariensis by WALLACH et al. 2014: 661.

Diagnosis: Sanzinia madagascariensis volontany differs from S. m. madagascariensis by a predominantly brownish adult colouration (predominantly greenish in S. m. m.) and a high genetic differentiation in the 16S rRNA gene (Tab. 1) which is nearly twice as high as between Acrantophis madagascariensis and A. dumerilii. Morphologically, there is a tendency of the new subspecies to have a lower number of infralabials, and possibly of supralabials and perioculars (see Tab. 3), but none of these characters can be used as a reliably diagnostic difference. 
EtymologyThe Malagasy word “volontany” means brown and refers to the predominant ground colour of the subspecies. It is used as invariable noun in apposition. 
References
  • BORA, PARFAIT; J. CHRISTIAN RANDRIANANTOANDRO, ROMA RANDRIANAVELONA, ELISOA F. HANTALALAINA, RAPHALI R. ANDRIANTSIMANARILAFY,<br />DANIEL RAKOTONDRAVONY, OLGA R. RAMILIJAONA, MIGUEL VENCES,<br />RICHARD K. B. JENKINS, FRANK GLAW, AND JÖRN KÖHLER 2009. AMPHIBIANS AND REPTILES OF THE TSINGY DE BEMARAHA PLATEAU, WESTERN MADAGASCAR: CHECKLIST, BIOGEOGRAPHY AND CONSERVATION. Herp. Cons. Biol. 5 (1):111-125.
  • Reynolds, R. Graham; Matthew L. Niemiller, Liam J. Revell 2014. Toward a Tree-of-Life for the boas and pythons: Multilocus species-level phylogeny with unprecedented taxon sampling. Molecular Phylogenetics and Evolution 71: 201–213 [published online in 2013] - get paper here
  • Vences, M. & F. Glaw 2003. Phylogeography, systematics and conservation status of boid snakes from Madagascar (Sanzinia and Acrantophis) Salamandra 39 (3-4): 181-206 - get paper here
 
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