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Scelarcis perspicillata (DUMÉRIL & BIBRON, 1839)

IUCN Red List - Scelarcis perspicillata - Least Concern, LC

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Higher TaxaLacertidae, Lacertinae, Sauria, Lacertoidea, Squamata (lizards)
SubspeciesScelarcis perspicillata chabanaudi (WERNER 1931)
Scelarcis perspicillata pellegrini (WERNER 1929)
Scelarcis perspicillata perspicillata (DUMÉRIL & BIBRON 1839) 
Common NamesE: Menorca Wall Lizard
G: Brilleneidechse
chabanaudi: Genetzte Brilleneidechse 
SynonymLacerta perspicillata DUMÉRIL & BIBRON 1839: 249
Lacerta perspicillata — GUICHENOT 1850: 12
Lacerta (Thetia) perspicillata — BOULENGER 1920: 335
Podarcis perspicillata — ARNOLD 1973
Lacerta perspicillata — WELCH 1982
Podarcis perspicillata — RICHTER 1986: 399
Podarcis perspicillata — ENGELMANN et al 1993
Teira perspicillata — MAYER & BISCHOFF 1996: 169
Scelarcis perspicillata — BONS & GENIEZ 1996
Podarcis perspicillata — SCHLEICH, KÄSTLE & KABISCH 1996: 441
Lacerta (Teira) perspicillata — HARRIS et al. 2003
Teira perspicillata — SCHLÜTER 2004: 44
Scelarcis perspicillata — ARNOLD et al. 2007
Teira perspicillata — KWET 2010
Scelarcis perspicillata — MEDIANI et al. 2015
Scelarcis perspicillata — LANUZA et al. 2016
Scelarcis perspicillata — MENDES et al. 2016

Scelarcis perspicillata chabanaudi (WERNER, 1931)
Lacerta perspicillata chabanaudi WERNER, 1931: 286
Lacerta perspicillata chabanaudi — HEDIGER 1935
Lacerta perspicillata chabanaudi — TIEDEMANN et al. 1994
Teira perspicillata chabanaudi — SCHLÜTER 2003
Lacerta (Scelarcis) perspicillata chabanaudi — SINDACO & JEREMČENKO 2008
Scelarcis perspicillata chabanaudi — PEREIRA et al. 2019

Scelarcis perspicillata pellegrini (WERNER, 1929)
Lacerta perspicillata pellegrini WERNER 1929: 5
Lacerta perspicillata pellegrini — WERNER 1931: 286
Lacerta perspicillata pellegrini forma Tazana WERNER 1931
Lacerta (Scelarcis) perspicillata pellegrini — SINDACO & JEREMČENKO 2008
Scelarcis perspicillata pellegrini — PEREIRA et al. 2019 
DistributionW/C Morocco (Middle and High Atlas), N/W Algeria, Spain (Menorca Island, introduced)

chabanaudi: Morocco; Type locality: Chella bei Rabat (= Chellah de Rabat); Fes, Marokko.

pellegrini: Morocco; Type locality: Safron (= Sefrou), Marokko;  
TypesHolotype: MNHN-RA 2749
Syntypes: NMW 16225, female; NMW 11153, female, MCZ 29981 [chabanaudi]
Holotype: MCZ R 27390 (29983 in catalogue); Syntype: NMW 11154; paratype: MNHN [pellegrini] 
DiagnosisDiagnosis (genus): The only genus in the Lacertini with a transparent window in the lower eyelid consisting of a single scale. Also possessing the following features found only in a minority of other Lacertini: usual number of presacral vertebrae 26–27 in males, medial loop of clavicle always interrupted posteriorly, lateral arms of interclavicle directed obliquely backwards, hemipenis with large sulcal lips on lobes, outer edge of parietal scale reaching lateral border of parietal table both posteriorly and anteriorly, nostril separated from first upper labial scale, five or more upper labial scales in front of subocular; no enlarged masseteric scale, 10–12 longitudinal rows of ventral scales, preanal scale often bordered by two semicircles of smaller scales, some animals without markings and others with broad dorsolateral stripes; no blue spots on lateral edges of the outer ventral scales. Other more widely distributed features include: head not strongly depressed, usually seven premaxillary teeth in adults, inscriptional ribs usually absent, tail brightly coloured in hatchlings, hemipenial microornamentation of hook-like spines (from Arnold et al. 2007: 55). 
CommentSubspecies: Besides the subspecies chabanaudi and pellegrini, Werner 1931 also described several “forms”, namely “Rerahiensis” and “Tazana”.

Genus: A separate (sub-) genus Scelarcis was proposed for this distinct species. BONS & GENIEZ (1996) reject the subspecies of P. perspicillata because of their mixed distribution pattern.

Type species: Lacerta perspicillata Duméril & Bibron, 1839 is the type species of the genus Scelarcis FITZINGER 1843.

Phylogenetics. Mitochondrial data (Harris and Arnold 1999, Oliverio et al. 2000) and morphology (Arnold 1989) suggest Teira perspicillata is a sister taxon to Teira dugesii (Milne-Edwards, 1829) which has been confirmed by Mendes et al. 2016. It is thus best placed in Teira (type species: dugesii) for the time being (although Mendes et al. 2016 maintain it in a separate genus). Preliminary evicence of large amount of genetic divergence concordant with sharp morphological variation (Harris et al. 2003) suggest that several species are presently confounded within this species (Jeroen Speybroeck, pers. comm. and cited references). 
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  • Barata, Mafalda; Ana Perera, D. James Harris, Arie van der Meijden, Salvador Carranza, Francisco Ceacero, Enrique García-Muñoz, Duarte Gonçalves, Sérgio Henriques, Fátima Jorge, Jonathan C. Marshall, Luis Pedrajas and Pedro Sousa 2011. New observations of amphibians and reptiles in Morocco, with a special emphasis on the eastern region. Herpetological Bulletin (116): 4 - get paper here
  • Berg, M.P. van den & Zawadzki, M. & Kroniger, M. 2013. Herpetological trip to Menorca (Balearics, Spain): April 30 till May 11, 2012. - get paper here
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  • Boulenger, George A. 1891. Catalogue of the reptiles and batrachians of Barbary (Morocco, Algeria, Tunisia), based chiefly upon the notes and collections made in 1880-1884 by M. Fernand Lataste. Tr. Zool. Soc. 13: 93-164
  • Boulenger,G.A. 1920. Monograph of the Lacertidae, vol. 1. British Museum (London) - get paper here
  • Crochet, Pierre Andre; Raphael Leblois, Julien Pierre Renoult 2015. New reptile records from Morocco and Western Sahara. Herpetology Notes 8: 583-588 - get paper here
  • Duméril, A. M. C. and G. Bibron. 1839. Erpétologie Générale on Histoire Naturelle Complète des Reptiles. Vol. 5. Roret/Fain et Thunot, Paris, 871 pp. - get paper here
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  • Pereira, Ana; Raquel Xavier, Ana Perera, Daniele Salvi, D James Harris 2019. DNA metabarcoding to assess diet partitioning and feeding strategies in generalist vertebrate predators: a case study on three syntopic lacertid lizards from Morocco. Biological Journal of the Linnean Society, Volume 127, Issue 4, August 2019, Pages 800–809, - get paper here
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