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Sceloporus heterolepis BOULENGER, 1895

IUCN Red List - Sceloporus heterolepis - Least Concern, LC
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Higher TaxaPhrynosomatidae, Sceloporinae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Dorsalkeel Spiny Lizard, Southern Odd-scaled Lizard
S: Lagartija de Escamas Dispares 
SynonymSceloporus heterolepis BOULENGER 1895
Sceloporus heterolepis — SMITH 1939: 197
Sceloporus heterolepis — SMITH & TAYLOR 1950: 121
Sceloporus heterolepis — LINER 1994
Sceloporus heterolepis — BELL et al. 2003
Sceloporus heterolepis — HEIMES 2022 
DistributionMexico (W Jalisco, Durango, Michoacan, Nayarit)

Type locality: La Cumbre de los Arrastrados (to which the type locality is restricted by SMITH & TAYLOR 1950), Real Alto “Riocho” [= Rancho ?] La Berberia, Sierra de Bolaños.  
Reproductionviviparous (ovoviviparous) 
TypesSyntypes: BMNH 1946.8.10.37-43 (formerly 1892.9.5.17-23), La Cumbre de los Arrastrados, Jalisco, 8500 ft.; 1946.8.10.25-30 (formerly 1892.10.31.4-9), La Cumbre de los Arrastrados, Jalisco, 8500 ft.; 1946.8.10.20 (formerly 1893.4.29.1-2), Real Alto, Jalisco, 7800 ft.; 1892.2.8.30, “Riocho (= Rancho)” La Berberia. All collected by A. C. Buller; MCZ 32346 
DiagnosisDiagnosis: Smith 1939: 197


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CommentSubspecies: Sceloporus heterolepis shannonorum LANGEBARTEL 1959 is now considered as a valid species. 
EtymologyThe Greek words heteros, "different," and lepido-, "scale," refer to the noticeably unequal sizes of the dorsal scales. 
References
  • Bell, E.L.; Smith, H.M. & Chiszar, D. 2003. AN ANNOTATED LIST OF THE SPECIES-GROUP NAMES APPLIED TO THE LIZARD GENUS SCELOPORUS. Acta Zoologica Mexicana (n.s.) 90: 103-174 - get paper here
  • Boulenger, G.A. 1895. Second report on additions to the lizard collection in the Natural History Museum. Proc. Zool. Soc. London 1894: 722-736 [published in April 1895 fide A. Bauer] - get paper here
  • Boulenger, G.A. 1897. A revision of the lizards of the genus Sceloporus. Proc. Zool. Soc. London 1897: 474-522 - get paper here
  • Cobarrubias, Edgar Flores; Daniel Cruz-Sáenz, and David Lazcano 2012. Notes on the Herpetofauna of Western Mexico 6: Amphibians and Reptiles of Hostotipaquillo, Jalisco, Mexico. Bull. Chicago Herp. Soc. 47 (2):21-26 - get paper here
  • Heimes, P. 2022. LIZARDS OF MEXICO - Part 1 Iguanian lizards. Edition Chimaira, Frankfurt Am Main, 448 pp. - get paper here
  • Köhler, G. & Heimes, P. 2002. Stachelleguane. Herpeton-Verlag (Offenbach), 174 pp.
  • Lara-Góngoro, Guillermo 2004. A new species of Sceloporus (Reptilia, Sauria: Phrynosomatidae) of the grammicus complex from Chihuahua and Sonora, Mexico. Bull. Maryland Herp. Soc. 40(1): 1-41 - get paper here
  • Leaché, A.D. 2010. Species trees for spiny lizards (Genus Sceloporus): Identifying points of concordance and conflict between nuclear and mitochondrial data. Molecular Phylogenetics and Evolution 54: 162–171 - get paper here
  • Leaché, A.D. & J.W.Sites, Jr. 2009. Chromosome Evolution and Diversification in North American Spiny Lizards (Genus Sceloporus). Cytogenet Genome Res 127: 166-181 - get paper here
  • Lemos-Espinal JA, Smith GR, Gadsden-Esparza H, Valdez-Lares R, Woolrich-Piña GA 2018. Amphibians and reptiles of the state of Durango, Mexico, with comparisons with adjoining states. ZooKeys 748: 65-87 - get paper here
  • Loc-Barragán JA, Smith GR, Woolrich-Piña GA, Lemos-Espinal JA 2024. An updated checklist of the amphibians and reptiles of Nayarit, Mexico with conservation status and comparison with adjoining States. Herpetozoa 37: 25-42 - get paper here
  • Loc-Barragán, Jesus, Juan Pablo Ramirez-Silva and Guillermo A. Woolrich-Piña. 2016. Geographic Distribution: Sceloporus heterolepis (Odd-scaled Spiny Lizard). Herpetological Review 47 (2): 260 - get paper here
  • Rojo-Gutiérrez, Juan Rubén; David Jaimes-Rodríguez , Daniel Cruz-Sáenz , Laura López-Fernández , Enrique Chávez-Uribe and David Lazcano 2018. Notes on the Herpetofauna of Western Mexico 19: An Update to the Herpetofauna of Volcán de Tequila in Jalisco, Mexico. Bulletin of the Chicago Herpetological Society 53(8):165-169 - get paper here
  • Rosas-Espinoza, V.C.; Rodríguez-Zaragoza, F.A.; Álvarez-Grzybowska, E.; Peña-Joya, K.E.; Santiago-Pérez, A.L.; Godoy-González, A.A.; Huerta-Martínez, F.M. 2024. Taxonomic and Functional Diversity of Reptiles in a Heterogeneous Landscape of Jalisco State, West-Central Mexico. Diversity 2024, 16, 394 - get paper here
  • Smith, H. M. 1939. The Mexican and Central American lizards of the genus Sceloporus. Field Mus. Nat. Hist., Zool. Ser. 26: 1-397. - get paper here
  • Smith, H.H.; Liner, E.A.; Ponce-Campos, P. & Chiszar, D. 2006. Sceloporus shannonorum LANGEBARTEL - Shannon’s Spiny Lizard. Catalogue of American Amphibians and Reptiles 813: 1-3 - get paper here
  • Smith, H.H.; Ponce-Campos, P.; Liner, E.A. & Chiszar, D. 2004. Sceloporus heterolepis Boulenger. Catalogue of American Amphibians and Reptiles 797: 1-3 - get paper here
  • Smith, H.M. & Taylor,E.H. 1950. An annotated checklist and key to the reptiles of Mexico exclusive of the snakes. Bull. US Natl. Mus. 199: 1-253 - get paper here
  • Webb, R.G. 1984. Herpetogeography in the Mazatlán-Durango Region of the Sierra Madre Occidental, Mexico. Vetrebrate Ecology and Systematics - A ribute to Henry S. Fitch; Museum of Natural History, University of Kansas, Lawrence, pp. 217-241
  • Woolrich-Piña, Guillermo A, Paulino Ponce-Campos, Jesús Loc-Barragán, Juan Pablo Ramírez-Silva, Vicente Mata-Silva, Jerry D. Johnson, Elí García-Padilla and Larry David Wilson. 2016. The herpetofauna of Nayarit, Mexico: composition, distribution, and conservation status. Mesoamerican Herpetology 3 (2): 376-448 - get paper here
 
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