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Sceloporus scalaris WIEGMANN, 1828

IUCN Red List - Sceloporus scalaris - Least Concern, LC
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Higher TaxaPhrynosomatidae, Sceloporinae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Light-bellied Bunch Grass Lizard
S: Lagartija Escamosa
E: Brown’s Bunchgrass Lizard [brownorum]
S: Escamosa de Brown [brownorum]
E: Southwestern Bunchgrass Lizard [unicanthalis]
S: Escamosa de Pastizal Sureña [unicanthalis] 
SynonymSceloporus scalaris WIEGMANN 1828: 370
Tropidolepis scalaris — DUMÉRIL & BIBRON 1837: 310
Tropidolepis Scalaris — DUMÉRIL, BIBRON & DUMÉRIL 1854: 271
Sceloporus scalaris — BOULENGER 1885: 234
Sceloporus scalaris scalaris — SMITH & TAYLOR 1950: 137
Sceloporus scalaris — STEBBINS 1985: 125
Sceloporus scalaris — LINER 1994
Sceloporus scalaris scalaris — BELL et al. 2003
Sceloporus scalaris — HEIMES 2022 
DistributionMexico (C Durango, southward over the central and southern parts of the Mexican Plateau; Distrito Federal, Guanajuato, Hidalgo, Jalisco, México, Michoacán, Puebla, Zacatecas, Nayarit, Aguascalientes, San Luis Potosí, Morelos, Querétaro, Veracruz, Tamaulipas).

Type locality: Mexico. Restricted to México, Distrito Federal, by SMITH & TAYLOR 1950.  
Reproductionoviparous 
TypesSyntypes: ZMB 654-656 (fide Taylor, 1969), collected by Ferdinand Deppe 
DiagnosisDiagnosis: Smith 1939: 338.


Additional details (33 characters) are available for collaborators and contributors. Please contact us for details. 
CommentTaxonomy: The S. scalaris species group includes five monotypic species, S. slevini, S. samcolemani, S. chaneyi, S. goldmani, and S. bicanthalis, and two polytypic species, S. scalaris (S. s. scalaris, S. s. unicanthalis, S. s. brownorum) and S. aeneus (S. a. aeneus, S. a. subniger).

Subspecies: Sceloporus scalaris brownorum, slevini, samcolemani and unicanthalis have been elevated to species status.

Distribution: reported from the US (Arizona), e.g. by Gloyd 1937. 
EtymologyNamed after the Latin scalaris, "of a ladder," refers to the successive transverse dorsal dark lines resembling a ladder. 
References
  • America, G. & Ortega, A. 1986. Mating behavior of the mountain lizard, Sceloporus scalaris. Bull. Maryland Herp. Soc. 22 (1): 23-25 - get paper here
  • Anderson, J.D. 1962. Egg laying and nesting in Sceloporus scalaris slevini Herpetologica 18 (3): 162-164. - get paper here
  • Anderson, James D. 1972. Pattern Polymorphism in the Bunch-Grass Lizard, Sceloporus scalaris slevini Journal of Herpetology 6 (1): 80 - get paper here
  • Ballinger, R. E.;Congdon, J. D. 1981. Population ecology and life history of a montane lizard (Sceloporus scalaris) in southeastern Arizona. Journal of Natural History 15: 213-222 - get paper here
  • Ballinger, R. E.;Congdon, J. D. 1996. Status of the bunch grass lizard, Sceloporus scalaris, in the Chiricahua Mountains of southeastern Arizona. Bull. Maryland Herp. Soc. 32: 67-69 - get paper here
  • Ballinger, R.E. & J.D. Congdon 1980. Food Resource Limitation of Body Growth Rates in Sceloporus scalaris (Sauria: Iguanidae) Copeia 1980 (4): 921-923. - get paper here
  • Bell, E.L.; Smith, H.M. & Chiszar, D. 2003. AN ANNOTATED LIST OF THE SPECIES-GROUP NAMES APPLIED TO THE LIZARD GENUS SCELOPORUS. Acta Zoologica Mexicana (n.s.) 90: 103-174 - get paper here
  • Benabib, Miriam; Kjer, Karl M.; Sites, Jack W., Jr 1997. Mitochondrial DNA sequence-based phylogeny and the evolution of viviparity in the Sceloporus scalaris group (Reptilia, Squamata). Evolution 51 (4): 1262-1275 - get paper here
  • Blackburn, D.G. 1993. STANDARDIZED CRITERIA FOR THE RECOGNITION OF REPRODUCTIVE MODES IN SQUAMATE REPTILES. Herpetologica 49 (1): 118-132 - get paper here
  • Bock, C.E., H.M. Smith & J.H. Bock 1990. The Effect of Livestock Grazing Upon Abundance of the Lizard, Sceloporus scalaris, in Southeastern Arizona Journal of Herpetology 24 (4): 445-446. - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Boulenger, G.A. 1897. A revision of the lizards of the genus Sceloporus. Proc. Zool. Soc. London 1897: 474-522 - get paper here
  • Bryson, Robert W.; Uri Omar García-Vázquez, Brett R. Riddle 2011. Relative roles of Neogene vicariance and Quaternary climate change on the historical diversification of bunchgrass lizards (Sceloporus scalaris group) in Mexico. Molecular Phylogenetics and Evolution 62 (1): 447-457 - get paper here
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  • Carbajal-Márquez, Rubén A., Christian M. García-Balderas, Gustavo E. Quintero-Díaz, Ana G. Pérez-Delgadillo and Raramuri Reyes-Ardit. 2013. Sceloporus scalaris (light-bellied bunchgrass lizard) mating behavior. Herpetological Review 44 (4): 683-684 - get paper here
  • Castro Franco, R. & M. G. Bustos-Zagal 2003. Lagartijas de Morelos, México: distribución, hábitat y conservación. Acta Zoológica Mexicana (n.s.) 88:123-142 - get paper here
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  • CRUZ-ELIZALDE, RACIEL; AURELIO RAMÍREZ-BAUTISTA, ABRAHAM LOZANO, JULIÁN A. VELASCO, PABLO OCTAVIO-AGUILAR & CHRISTIAN BERRIOZABAL-ISLAS. 2022. Variation in size and shape sexual dimorphism in the Sceloporus scalaris species group (Squamata: Phrynosomatidae) from the Transvolcanic Belt of Mexico. Biological Journal of the Linnean Society 135(3): 499–517. - get paper here
  • Degenhardt, William G.; C. W. Painter, and A. H. Price 1996. Amphibians and reptiles of New Mexico. Univ. New Mexico Press, 431 pp.
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  • Fernández-Badillo, Leonardo & Irene Goyenechea-Mayer Goyenechea 2010. Anfibios y reptiles del valle del Mezquital, Hidalgo, México. Revista Mexicana de Biodiversidad 81: 705- 712
  • Fitch, H. S. 1978. Sexual size differences in the genus Sceloporus. Univ. Kansas Sci. Bull. 51 (13): 441-461 - get paper here
  • Gloyd, Howard K. 1937. A herpetological consideration of faunal areas in Southern Arizona. Bulletin of the Chicago Academy of Sciences 5 (5): 77-136 - get paper here
  • Gravenhorst, J. L. C. 1838. Beiträge zur genaueren Kenntniss einiger Eidechsengattungen. Nova Acta Acad. Caes. Leop.-Carol. 18: 712-784 [1837]
  • GRUMMER, JARED A. & ROBERT W. BRYSON JR. 2014. A new species of bunchgrass lizard (Squamata: Phrynosomatidae) from the southern sky islands of the Sierra Madre Occidental, Mexico. Zootaxa 3790 (3): 439–450 - get paper here
  • Grummer, Jared A.; Rob W. Bryson, Jr., and Tod W. Reeder 2014. Species Delimitation Using Bayes Factors: Simulations and Application to the Sceloporus scalaris Species Group (Squamata: Phrynosomatidae). Systematic Biology 63 (2): 119-133<br>doi:10.1093/sysbio/syt069 - get paper here
  • Heimes, P. 2022. LIZARDS OF MEXICO - Part 1 Iguanian lizards. Edition Chimaira, Frankfurt Am Main, 448 pp. - get paper here
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  • Muñoz-Nolasco, Francisco Javier; Daniel Cruz-Sáenz, Oscar Jair Rodríguez-Ruvalcaba and Isaac Enrique Terrones-Ferreiro 2015. Notes on the Herpetofauna of Western Mexico 12: Herpetofauna of a Temperate Forest in Mazamitla, Southeastern Jalisco, Mexico. Bull. Chicago Herp. Soc. 50 (4):45-50 - get paper here
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