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Sceloporus bimaculosus PHELAN & BRATTSTROM, 1955

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Higher TaxaPhrynosomatidae, Sceloporinae; Iguania, Sauria, Squamata (lizards) 
Subspecies 
Common NamesE: Twin-spotted Spiny Lizard 
SynonymSceloporus magister bimaculosus PHELAN & BRATTSTROM 1955
Sceloporus magister bimaculosus — CONANT & COLLINS 1991: 105
Sceloporus magister bimaculosus — LINER et al. 1993
Sceloporus magister bimaculosus — BELL et al. 2003
Sceloporus bimaculosus — SCHULTE et al. 2006
Sceloporus bimaculosus — LEMOS-ESPINAL & SMITH 2007
Sceloporus bimaculosus — CROTHER et al. 2012
Sceloporus bimaculosus — CROTHER et al. 2017 
DistributionUSA (New Mexico, Texas), Mexico (Coahuila, Chihuahua, Durango).

Type locality: "6.6 miles east of San Antonio, Socorro, New Mexico").  
Reproduction 
TypesHolotype: CAS 91199 
Diagnosis 
CommentHistorically, S. magister, including magister bimaculosus may have been confused with S. clarkii (STEJNEGER 1893).

Synonymy: Sceloporus magister bimaculosus was elevated to full species by Schulte et al. 2006) but returned to the synonymy of S. magister by Leaché and Mulcahy (2007). Recent authors have alternatively treated it as full species or synonym (e.g. Laczano et al. 2019 in whose list it does not occur in Coahuila). 
EtymologyThe name S. m. bimaculosus is a Latin adjective based upon bis-, "two" or "twice," and maculosus, "spotted," and refers to a series of paired brown blotches on dorsum. 
References
  • Bell, E.L.; Smith, H.M. & Chiszar, D. 2003. AN ANNOTATED LIST OF THE SPECIES-GROUP NAMES APPLIED TO THE LIZARD GENUS SCELOPORUS. Acta Zoologica Mexicana (n.s.) 90: 103-174 - get paper here
  • Burré, G. 2002. Begegnungen mit Iguaniden im Südwesten der USA. Iguana Rundschreiben 15 (2): 13-21
  • Burt, Charles E. 1933. Some lizards from the Great Basin of the West and adjacent areas, with comments on the status of various forms. American Midland Naturalist 14: 228-250 - get paper here
  • Conant,R. & Collins,J.T. 1991. A Field Guide to Reptiles and Amphibians of Eastern/Central North America, 3rd ed. Houghton Mifflin (Boston/New York), xx + 450 p.
  • Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92
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  • Eaton, Theodore H., Jr 1935. Report on amphibians and reptiles of the Navajo country. Bulletin 3. Rainbow Bridge-Monument Valley Expedition : 1-20
  • Fitch, H. S. 1978. Sexual size differences in the genus Sceloporus. Univ. Kansas Sci. Bull. 51 (13): 441-461 - get paper here
  • FLESCH, AARON D.; DON E. SWANN, DALE S. TURNER, AND BRIAN F. POWELL 2010. HERPETOFAUNA OF THE RINCON MOUNTAINS, ARIZONA. Southwestern Naturalist 55(2):240–253 - get paper here
  • Grismer, L. L. & J. A. MCGUIRE 1996. Taxonomy and biogeography of the Sceloporus magister complex (Squamata: Phrynosomatidae) in Baja California, México. Herpetologica 52 (3): 416-427. - get paper here
  • Hallowell,E. 1854. Description of new reptiles from California. Proc. Acad. Nat. Sci. Philad. 7 [1854]: 91-97 - get paper here
  • Hernandez T, Herr MW, Stevens S, Cork K, Medina-Nava C, Vialpando CJ, Warfel T, Fields N, Brodie C, Graham SP 2019. New distribution records for amphibians and reptiles in eastern Chihuahua, Mexico. Check List 15(1): 79-86 - get paper here
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  • Leaché, A.D. 2010. Species trees for spiny lizards (Genus Sceloporus): Identifying points of concordance and conflict between nuclear and mitochondrial data. Molecular Phylogenetics and Evolution 54: 162–171 - get paper here
  • Leaché, A.D. & J.W.Sites, Jr. 2009. Chromosome Evolution and Diversification in North American Spiny Lizards (Genus Sceloporus). Cytogenet Genome Res 127: 166-181 - get paper here
  • Leaché, A.D. & Mulcahy, D.G. 2007. Phylogeny, divergence times and species limits of spiny lizards (Sceloporus magister species group) in western North American deserts and Baja California. Molecular Ecology 16: 5216–5233 - get paper here
  • Lemos-Espinal JA, Smith GR, Gadsden-Esparza H, Valdez-Lares R, Woolrich-Piña GA 2018. Amphibians and reptiles of the state of Durango, Mexico, with comparisons with adjoining states. ZooKeys 748: 65-87 - get paper here
  • Lemos-Espinal, J.A. & Smith, H.M. 2007. Amphibians and reptiles of the state of Coahuila, Mexico. Universidad Nacional Autonoma de Mexico, 550 pp.
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  • Parker, W. S.;Pianka, E. R. 1973. Notes on the ecology of the Iguanid lizard, Sceloporus magister. Herpetologica 29 (2): 143-152 - get paper here
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  • Schulte II, James A.; J. Robert Macey and Theodore J. Papenfuss 2006. A genetic perspective on the geographic association of taxa among arid North American lizards of the Sceloporus magister complex (Squamata: Iguanidae: Phrynosomatinae). Molecular Phylogenetics and Evolution 39 (3): 873-880 - get paper here
  • Smith, H. M. 1939. The Mexican and Central American lizards of the genus Sceloporus. Field Mus. Nat. Hist., Zool. Ser. 26: 1-397. - get paper here
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  • Tanner, Vasco M.;Hayward, C. Lynn 1934. A biological study of the La Sal Mountains, Utah report No. 1 (Ecology). Proceedings of the Utah Academy of Sciences, Arts, and Letters 11: 209-235
  • Tanner, W. W.;Krogh, J. E. 1973. Ecology of Sceloporus magister at the Nevada test site, Nye County, Nevada. Great Basin Naturalist 33 (3): 133-146 - get paper here
  • Tanner, Wilmer W. 1955. A new Sceloporus magister from eastern Utah. Great Basin Naturalist. 15: 32-34 - get paper here
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  • Taylor, Harry L.;Hollard, Richard L.;Humphrey, Rosanna L.;Smith, Hobart M. 1996. Geographic Distribution. Sceloporus magister cephaloflavus. Herpetological Review 27 (2): 87-88 - get paper here
  • Tevis, lloyd, Jr. 1944. Herpetological notes from lower California. Copeia 1944 (1): 6-18 - get paper here
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  • Turner, Frederick B.;Wauer, Roland H. 1963. A survey of the herpetofauna of the Death Valley area. Great Basin Naturalist 23 (3-4): 119-128 - get paper here
  • Valdez-Lares, R.; R. Muñiz-Martínez; E.Gadsden; G. Aguirre-León; G. Castañeda-Gaytán; R. Gonzalez-Trápaga 2013. Checklist of amphibians and reptiles of the state of Durango, México. Check List 9 (4):714-724 - get paper here
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