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Sceloporus binocularis DUNN, 1936

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Higher TaxaPhrynosomatidae, Sceloporinae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Nuevo Leon Crevice Swift
S: Espinosa de Montafia 
SynonymSceloporus binocularis DUNN 1936: 474
Sceloporus ferrariperezi binocularis — SMITH 1939: 216
Sceloporus torquatus binocularis — SMITH & TAYLOR 1950: 126
Sceloporus torquatus binocularis — BELL et al. 2003
Sceloporus torquatus binocularis — LINER & CASAS-ANDREU 2008
Sceloporus binocularis — CAMPILLO-GARCÍA et al. 2021 
DistributionMexico (Nuevo León)

Type locality: Trail from Pablillo to Alamar, Nuevo León.  
Reproductionviviparous 
TypesHolotype: ANSP 20032, collected by H.A. Pilsbry, F.W. Pennell and C.H. Harvey Malnate in 1934. 
DiagnosisDiagnosis (binocularis): A Sceloporus with rather uniform body scalation of large scales; dorsal rows parallel, lateral oblique; supraorbital region occupied by a single row of enlarged scales; femoral pores in widely separated rows, 11-13; no black neck collar; no definite markings save two light spots on nape (Dunn 1936: 474).

Description (binocularis): Head scales smooth; frontal transversely divided (anterior frontal longitudinally divided in 20019); two canthal scales; nasal sep- arated from rostral by a row of scales bordering rostral and labials; four small internasals, quite unsymmetrical; two rows of scales between labials and suboculars; four or five postmental pairs, anterior in contact medially; four large supraocular scales, bordered internally by a row of granules, separated from supraciliaries by one (in spots two) row of granules; fronto- parietals separate except in 20019; two parietals, anterior largest (three on right side of 20032, second largest); three auricular denticles of medium size, two upper largest; dorsal body scales larger than ventrals, lateral intermediate; dorsals parallel, keeled, weakly mucronate; laterals oblique, keeled; ventrals smooth; 23-25 dorsals between head and base of legs; 7-9 in head length; cervical fold weak, all scales large; ventral scales of thigh flat, of tibia keeled; femoral pores in rows separated by 5 scales; 11-13, the number eleven on three legs, twelve on two, thirteen on one; fourth toe measured from base of fifth equals snout to anterior border of ear; snout to vent of largest specimen (type) 56 mm.; color in general greenish, spotted irregularly with single dark scales; throat mottled light blue and white; head dark above; head color extends onto nape and suprascapular area, but is interrupted by two longitudinal light spots or bars on nape; belly light (Dunn 1936: 474).

Remarks (binocularis): A Sceloporus with very large body scales, with large supra- ocular scales, and lacking a black collar, seems very unusual when one investigates the literature or examines specimens. The only Sceloporus hitherto described from the plateau of Mexico, which conforms with these criteria, is S. viviparus, but its detailed description certainly does not fit the present form. In the absence of other evidence it may be regarded as an ally (Dunn 1936: 475).

See also detailed Diagnosis and description by Smith 1939: 216. 
CommentSympatry: Circular sympatry may occur between S. t. mikeprestoni, derived from S. t. torquatus, and S. t. binocularis, derived from S. t. melanogaster. Additional field work in the hiatus between the ranges of S. t. mikeprestoni and S. t. binocularis is necessary to resolve this problem (BELL et al. 2003) but see CAMPILLO-GARCÍA et al. 2021 for an update.

Group: S. torquatus group (sensu lato, fide Flores-Villela et al. 2022) 
EtymologyNamed after the Latin adjective from “bis-” = two or paired, and “oculus” = eye, referring to the two enlarged supraocular scales. 
References
  • Bell, E.L.; Smith, H.M. & Chiszar, D. 2003. AN ANNOTATED LIST OF THE SPECIES-GROUP NAMES APPLIED TO THE LIZARD GENUS SCELOPORUS. Acta Zoologica Mexicana (n.s.) 90: 103-174 - get paper here
  • Campillo-García G, Flores-Villela O, Butler BO, Velasco Vinasco JA, Ramírez Corona F 2021. Hidden diversity within a polytypic species: The enigmatic Sceloporus torquatus Wiegmann, 1828 (Reptilia, Squamata, Phrynosomatidae. Vertebrate Zoology 71: 781-798 - get paper here
  • Dunn, Emmett Reid 1936. The amphibians and reptiles of the Mexican Expedition of 1934. Proceedings of the Academy of Natural Sciences of Philadelphia 88: 471-477 - get paper here
  • FLORES-VILLELA, Oscar, SMITH, E. N., CAMPILLO-GARCÍA, Gustavo, MARTÍNEZ-MÉNDEZ, Norberto, & CAMPBELL, J. A. 2022. A new species of Sceloporus of the torquatus group (Reptilia: Phrynosomatidae) from West Mexico. Zootaxa 5134 (2): 286-296 - get paper here
  • Liner, Ernest A., and Gustavo Casas-Andreu. 2008. Standard Spanish, English and Scientific Names of the Amphibians and Reptiles of Mexico. Herpetological Circular 38: 167 p.
  • Smith, H. M. 1939. The Mexican and Central American lizards of the genus Sceloporus. Field Mus. Nat. Hist., Zool. Ser. 26: 1-397. - get paper here
 
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