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Sceloporus merriami STEJNEGER, 1904

IUCN Red List - Sceloporus merriami - Least Concern, LC

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Higher TaxaPhrynosomatidae, Sceloporinae; Iguania, Sauria, Squamata (lizards) 
SubspeciesSceloporus merriami annulatus SMITH 1937
Sceloporus merriami australis WILLIAMS, SMITH & CHRAPLIWY 1960
Sceloporus merriami ballingeri LEMOS-ESPINAL, SMITH, AUTH & CHISZAR 2001
Sceloporus merriami longipunctatus OLSON 1973
Sceloporus merriami merriami STEJNEGER 1904
Sceloporus merriami sanojae LEMOS-ESPINAL in SMITH, LEMOS-ESPINAL & CHISZAR 2003
Sceloporus merriami williamsi LEMOS-ESPINAL, CHISZAR & SMITH 2000 
Common NamesE: Canyon Lizard
S: Lagartija de Cañón
annulatus: Big Bend Canyon Lizard
australis: Southeastern Canyon Lizard
ballingeri: Ballinger’s Canyon Lizard
longipunctatus: Presidio Canyon Lizard
merriami: Merriam's Canyon Lizard
sanojae: Sanoja's Canyon Lizard
williamsi: Williams’ Canyon Lizard 
SynonymSceloporus merriami STEJNEGER 1904
Sceloporus merriami merriami — SMITH & TAYLOR 1950: 131
Sceloporus merriami — CONANT & COLLINS 1991: 109
Sceloporus merriami — LINER 1994
Sceloporus merriami merriami — BELL et al. 2003
Sceloporus merriami merriami — CROTHER et al. 2012

Sceloporus merriami annulatus SMITH 1937: 83
Sceloporus merriami annulatus — BARTLETT & BARTLETT 1999: 225
Sceloporus merriami annulatus — BELL et al. 2003
Sceloporus merriami annulatus — LINER & CASAS-ANDREU 2008
Sceloporus merriami annulatus — CROTHER et al. 2012

Sceloporus merriami australis WILLIAMS, SMITH & CHRAPLIWY 1960
Sceloporus merriami australis — BELL et al. 2003
Sceloporus merriami australis — LINER & CASAS-ANDREU 2008

Sceloporus merriami ballingeri LEMOS-ESPINAL, SMITH, AUTH & CHISZAR 2001
Sceloporus merriami ballingeri — LINER & CASAS-ANDREU 2008

Sceloporus merriami longipunctatus OLSON 1973
Sceloporus merriami longipunctatus — BARTLETT & BARTLETT 1999: 227
Sceloporus merriami longipunctatus — BELL et al. 2003
Sceloporus merriami longipunctatus — CROTHER et al. 2012

Sceloporus merriami sanojae LEMOS-ESPINAL in SMITH et al. 2003
Sceloporus merriami sanojae — BELL et al. 2003
Sceloporus merriami sanojae — LINER & CASAS-ANDREU 2008

Sceloporus merriami williamsi LEMOS-ESPINAL, CHISZAR & SMITH 2000
Sceloporus merriami williamsi — BELL et al. 2003
Sceloporus merriami williamsi — LINER & CASAS-ANDREU 2008 
DistributionUSA (SW Texas), Mexico (Nuevo Léon, Coahuila, Chihuahua)

merriami: Rio Grande and its immediate tributaries from western Brewster County to SE Val Verde County, Texas; Mexico (Chihuahua [HR 32: 276]). Type locality: East Painted Cave, near mouth of the Pecos River, Texas.

annulatus: S/C Brewster County, Texas. Mexico (E Coahuila). Type locality: East slope of the Chisos Mountains, Brewster County, Texas.

australis: Mexico (Coahuila). Type Locality: 15.6 mi (25.1 km) E Cuatro Ciénegas, Coahuila, Mexico.

ballingeri: Mexico (Durango). Type Locality: La Campana, Sierra La Campana, municipio Tlahualilo, Durango (26°7'39.1"N, 103°41'00.0"W).

longipunctatus: USA (W Texas); Type Locality: Closed Canyon, 14.3 rd mi (23.0 km.) SE Redford, Presidio County, Texas.

merriami: W Texas, Mexico (Coahuila)

sanojae: Mexico (Chihuahua). Type Locality: Rancho Peñoles (27°7'49.6"N, 103°48'45.0"W), 1194m, Chihuahua, Mexico.

williamsi: Mexico (Chihuahua). Type Locality: El Fortín, 51 air km W Rio Grande at 29°35’40”N. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesHolotype: USNM 33039
Holotype: UIMNH 25058, EHT-HMS A787 [annulatus]
Holotype: UIMNH 43319, collected 9 July 1958, by P. S. [australis]
Holotype: UBIPRO 5094, Julio A. Lemos-Espinal collector, 9 June 2000 [ballengeri]
Holotype: USNM 192744, formerly REO 3286, collected 31 May 1971, by R. Earl Olson [longipunctatus]
Holotype: UBIPRO 7456, collected 24 July 2001 by Julio A. Lemos-Espinal [sanojae].
Holotype: UIMNH 52378, taken by K. L. Williams, E. O. Moll, F. Vuilleumier, and J. E. Williams,
31 August 1962 [williamsi] 
Comment 
EtymologyEtymology: This form honors C. Hart Merriam, the originator of the life-zone theory of biotic ditribution, and chief of the Bureau of Biological Survey, predecessor of the U. S. Fish and Wildlife Service.

Etymology: S. m. annulatus is named after the Latin noun annulus refers to the banded subcaudal surface, such banding being one of the key characteristics separating this subspecies from Sceloporus merriami merriami.

Etymology: S. m. australis has been named after the Latin adjective australis, "southern," referring to the range of this southernmost subspecies of S. merriami.

Etymology: S. m. ballingeri honors Dr. Royce E. Ballinger, of the University of Nebraska, for his many contributions to herpetology and for guidance of Julio A. Lemos-Espinal to his doctoral degree.

Etymology (longipunctatus): The Latin adjective long, "long," the noun punctum, "spot," and the suffix -atus, "pertaining to", refer to laterally elongate paravertebral spots, diagnostic of the subspecies.

Etymology (sanojae): The subspecific matronym honors Susy Sanoja Sarabia, in honor of and gratitude for her long-term support of Julio A. Lemos-Espinal in the field, laboratory and at home.

Etymology (williamsi): The patronym honors Kenneth L. Williams, long a specialist in the classification of snakes and the herpetology of Mexico and Honduras. In the original description, two spellings of the name appeared: willaimsi and williamsi. BELL et al. (2003) designated the latter as correct. 
References
  • Axtell, Ralph W. 1939. AMPHIBIANS AND REPTILES OF THE BLACK GAP WILDLIFE MANAGEMENT AREA, BREWSTER COUNTY, TEXAS. Southwestern Naturalist 4 (2): 88-109
  • Bartlett, R. D. & Bartlett, P. 1999. A Field Guide to Texas Reptiles and Amphibians. Gulf Publishing Co., Houston, Texas, 331 pp.
  • Bell, E.L.; Smith, H.M. & Chiszar, D. 2003. AN ANNOTATED LIST OF THE SPECIES-GROUP NAMES APPLIED TO THE LIZARD GENUS SCELOPORUS. Acta Zoologica Mexicana (n.s.) 90: 103-174
  • Chaney, Allan H.;Gordon, Robert E. 1954. Notes on a population of Sceloporus merriami merriami Stejneger. Texas Journal of Science 6 (1): 78-82
  • Cole, Charles J. 1971. Karyotypes of five monotypic species groups of lizards in the genus Sceloporus. American Museum Novitates (2450): 1-17 - get paper here
  • Conant,R. & Collins,J.T. 1991. A Field Guide to Reptiles and Amphibians of Eastern/Central North America, 3rd ed. Houghton Mifflin (Boston/New York), xx + 450 p.
  • Cooper Jr., W.E. 2005. Duration of movement as a lizard foraging movement variable. Herpetologica 61 (4): 363-372 - get paper here
  • Cooper Jr., W.E.; Vitt, L.J.; Caldwell, J.P. & Fox, S.F. 2005. Relationships among foraging variables, phylogeny, and foraging modes, with new data for nine North American lizard species. Herpetologica 61 (3): 250-259 - get paper here
  • Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92
  • De Los Reyes, M. N. and D. Lazcano 2016. Geographic Distribution: Sceloporus merriami (Canyon Lizard). Herpetological Review 47(3): 424.
  • Dixon, James R. 2000. Amphibians and reptiles of Texas, second edition. Texas A&M University Press, 421 pp.
  • Fitch, H. S. 1978. Sexual size differences in the genus Sceloporus. Univ. Kansas Sci. Bull. 51 (13): 441-461 - get paper here
  • Jones, L.L. & Lovich, R.E. 2009. Lizards of the American Southwest. A photographic field guide. Rio Nuevo Publishers, Tucson, AZ, 568 pp. [review in Reptilia 86: 84] - get paper here
  • Leaché, A.D. 2010. Species trees for spiny lizards (Genus Sceloporus): Identifying points of concordance and conflict between nuclear and mitochondrial data. Molecular Phylogenetics and Evolution 54: 162–171 - get paper here
  • Leaché, A.D. & J.W.Sites, Jr. 2009. Chromosome Evolution and Diversification in North American Spiny Lizards (Genus Sceloporus). Cytogenet Genome Res 127: 166-181 - get paper here
  • Lemos-Espinal JA, Smith GR, Cruz A 2016. Amphibians and Reptiles of the state of Nuevo León, Mexico. ZooKeys 594: 123-141, doi: 10.3897/zookeys.594.8289 - get paper here
  • Lemos-Espinal, J.A. & Smith, H.M. 2007. Amphibians and reptiles of the state of Chihuahua, Mexico. Universidad Nacional Autonoma de Mexico, 613 pp.
  • Lemos-Espinal, J.A. & Smith, H.M. 2007. Amphibians and reptiles of the state of Coahuila, Mexico. Universidad Nacional Autonoma de Mexico, 550 pp.
  • Lemos-Espinal, J.A.; Smith, H.M.; Auth, D.L. & Chiszar, D. 2001. The subspecies of Sceloporus merriami (Reptilia: Lacertilia) in Chihuahua and Durango, Mexico. Bull. Maryland Herp. Soc. 37 (4): 123-129
  • Lemos-Espinal, Julio A., David Chiszar, and Hobart M. Smith 2000. The lizard Sceloporus merriami in Chihuahua, Mexico. Bull. Maryland Herp. Soc. 36 (3): 86-96
  • Liner, Ernest A.;Montanucci, Richard R.;González-Alonso, Arturo;Mendoza Quijano, Fernando 1993. An additional contribution to the herpetology of Northern Coahuila, Mexico. Boletín de la Sociedad Herpetológica Mexicana 5 (1): 9-11 - get paper here
  • Lutterschmidt, William I. 2013. REPTILES IN RESEARCH: INVESTIGATIONS OF ECOLOGY, PHYSIOLOGY, AND BEHAVIOR FROM DESERT TO SEA. Nova Science, New York, 527 pp. - get paper here
  • Olson R E 1979. Sceloporus merriami Stejneger. Canyon lizard. Catalogue of American Amphibians and Reptiles (227): 1-2 - get paper here
  • Olson, R. Earl 1973. Variation in the Canyon lizard, Sceloporus merriami Stejneger. Herpetologica 29 (2): 116-127 - get paper here
  • Smith, H. M. 1939. The Mexican and Central American lizards of the genus Sceloporus. Field Mus. Nat. Hist., Zool. Ser. 26: 1-397. - get paper here
  • Smith, H. M.; Lemos-Espinal, J. A.; Chiszar, D. 2003. New subspecies of Sceloporus merriami (Reptilia: Lacertilia) and the derivation of its subspecies. Southwestern Naturalis 48 (4): 700-704 - get paper here
  • Smith,H.M. 1937. A new subspecies of the lizard genus Sceloporus from Texas. Proc. Biol. Soc. Washington 50: 83-86 - get paper here
  • Stejneger,L. 1904. A new lizard from the Rio Grande Valley, Texas. Proc. Biol. Soc. Washington. 17: 17-20 - get paper here
  • Valdez-Lares, R.; R. Muñiz-Martínez; E.Gadsden; G. Aguirre-León; G. Castañeda-Gaytán; R. Gonzalez-Trápaga 2013. Checklist of amphibians and reptiles of the state of Durango, México. Check List 9 (4):714-724 - get paper here
  • Werning, Heiko 2012. Die Reptilien und Amphibien des Südwestens. Draco 13 (50): 18-60 - get paper here
  • Wright, A. H.;Wright, A. A. 1927. Notes on Sceloporus merriami Stejneger. Proc. Biol. Soc. Washington 40: 57-64 + 3 plates - get paper here
 
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